A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data Author Tarasov, Sergei text Zootaxa 2017 2017-10-04 4329 2 101 149 journal article 31902 10.11646/zootaxa.4329.2.1 7e33ec7d-211b-4042-8d07-344364ee805d 1175-5326 1001915 8226E27D-E3A9-481A-B51B-558643Bbb03A Parachorius newthayerae Tarasov new species ( Figs. 17 , 26F ) urn:lsid:zoobank.org:act:5DE8A2B0-44BC-43A0-B38F-B3483509F42F Type locality . Bolaven plateau, Laos. Etymology. The species is named after two great entomologists, Alfred Newton and Margaret Thayer, who collected the largest series of specimens of this new species in Laos . Alfred and Margaret kindly hosted me during a visit to the Field Museum where the largest part of this taxonomic work on Parachorius was conducted. The epithet “ newthayerae ” is the combination of their last names. This species name should be treated as a noun in apposition. Distribution. This species seems endemic to Bolaven plateau in southern Laos . Taxonomic notes. In females of P. newthayerae the secondary sexual characters of legs modification are not expressed, although anterior angles of pronotum can be very slightly angulate. As only the secondary sexual characters reliably separate P. newthayerae from P. maruyamai , P. fingorum , and P. thomsoni , the identification of P. newthayerae using females is ambiguous. Diagnosis. This species is most closely related to P. maruyamai . Both species differ from all other Parachorius by sharped frontal angles of pronotum along with strongly curved metatibia in males. Parachorius newthayerae differs from P. maruyamai by dilated posterior margin of metafemur and angulate lateral margin of pronotum. Description. Body. Body black, length 7.8–8.0 mm. Elytra uniformly colored. Head. Clypeus with two indistinct teeth, outer margin of each tooth not notched basally. Pronotum. Pronotum distinctly angulate laterally. Frontal angles sharp. Disc covered with punctures separated by 0.5–2.0 puncture diameter(s). Pronotum in anterior-posterior direction notably shorter than elytra. Elytra. Elytra covered with sparse punctures. Wings. Wings normally developed. Legs. Profemur with almost straight or sinuate anterior margin. Protibia with apical spur approximately reaching middle of protarsomere 3. First tooth almost not modified in males, approximately as wide as second tooth; first and second teeth distinctly separated from each other; first tooth normal, distinctly separated from denticles located between first and second teeth. Metafemoral posterior margin serrate (sometimes slightly) or not serrate, medially dilated, and produced in large rounded angle. Metatibia strongly curved and notched medially; inner margin not denticulate, produced in small angle apically. Metasternum. Transverse basal ridge of metasternum long, at least 1/5 of mesocoxa length. Aedeagus. Parameres symmetrical, simple, apex widely rounded and inferiorly angulate. Holotype . Male. Profemur with sinuate anterior margin. Metafemoral posterior margin serrate. FIGURE 17. Morphology of Parachorius newthayerae . (A) aedeagus lateral view (paratype). (B) aedeagus lateral view. (C) habitus (male, paratype). (D) left metafemur (male, paratype). (E, G) propleuron (female). (F) propleuron (male, paratype). Material examined. Holotype : Laos : 1 male , " LAOS : Champasak Pr.: Bolaven Plateau, Ban Thôngvay (= Xékatam ) vic., old logging road N of village 1095m , 15°14.288'N 106°31.891'E , 8-16.vi.2008 / selectively logged forest; FMHD#2008-037 , flight intercept trap , A. Newton & M. Thayer ; ANMT site 1231 ex 95% FIELD MUS. NAT. HIST. / HOLOTYPUS Parachorius newthayerae / S. Tarasov det. 2011", 15°14.288'N, 106°31.899'E (FMNH). Paratypes (3 males, 3 females) : 1 male paratype , Champasak Province : Bolaven Plateau, Ban Thôngvay (= Xékatam ) vicinity, old logging road N of village, FMHD#2008-034 , ANMT site 1230, 15°13.96'N, 106°31.731'E, 1035 m , selectively logged forest, on 8–16/vi/2008 (A. Newton, M. Thayer) (FMNH); 1 male paratype , Champasak Province : Bolaven Plateau, Ban Thôngvay (= Xékatam ) vicinity, old logging road N of village, FMHD#2008-037 , ANMT site 1231, 15°14.288'N, 106°31.899'E, 1095 m , selectively logged forest, on 8– 16/vi/2008 (A. Newton, M. Thayer) (FMNH); 1 female paratype , Champasak Province : Bolaven Plateau, Ban Thôngvay (= Xékatam ) vicinity, old logging road N of village, FMHD#2008-038 , ANMT site 1231, 15°14.288'N, 106°31.891'E, 1095 m , selectively logged forest, on 8–16/vi/2008 (A. Solodovnikov, M. Thayer, A. Newton) (FMNH); 2 female paratypes , Champasak Province : Bolaven Plateau, Ban Thôngvay (= Xékatam ) vicinity, old logging road N of village, FMHD#2008-040 , ANMT site 1232, 15°14.494'N, 106°31.807'E, 1170 m , selectively logged forest, on 8–16/vi/2008 ( A. Newton , M. Thayer ) ( FMNH ); 1 male paratype , Champasak Province , Bolaven Plateau , Ban Thongvay , 15°14.741'N , 106°31.916'E , 1150 m , disturbed montane forest ( S. Tarasov ) (CST). Additional specimen: 1 female , Champasak Province : Bolaven Plateau, Ban Thôngvay (= Xékatam ) vicinity, old logging road N of village, FMHD #2008 -035, ANMT site 1230, 15°13.96'N, 106°31.731'E, 1035 m , selectively logged forest, on 8–16/vi/2008 (A. Solodovnikov, M. Thayer, A. Newton) (FMNH).