A Taxonomic Study Of The Myosoma Genus-Group With Description Of Amyosoma Cavei Sp. N. From Honduras (Hymenoptera: Braconidae: Braconinae: Braconini)
Author
Papp, J.
text
Acta Zoologica Academiae Scientiarum Hungaricae
2012
2012-05-10
58
1
1
29
journal article
106153
10.5281/zenodo.5732026
508a28c5-cba2-4fab-91a6-57b0cab1c01a
2064-2474
5732026
THE
MYOSOMA
GENUS–GROUP
Within the tribe
Braconini
the
Myosoma
genus-group is distinguished by the following features: 1.) First tergite very long and narrow, two to five times as long as broad behind (
Fig.
83
in QUICKE 1997: 172;
Figs 29
,
46
,
56
) 2.) Second tergite considerably transverse, i.e. 3.3–4 times as broad behind as long medially, laterally narrowing (
Figs 37
,
46
,
65
,
81
,
91
,
97
). 3.) Second submarginal cell long, i.e.
3–SR
at least 1.5–1.6 times, frequently more than twice, longer than
2–SR
(
Figs 26
,
34
,
42
,
53
,
62
,
71
,
79
,
89
,
95
).
The genus-group comprises three genera:
Amyosoma
VIERECK
,
Myosoma
BRULLÉ and
Mysomatoides
QUICKE. The
three genera and the species of
Myosoma
and
Myosomatoides
are separated by the features keyed:
1 (4) Second tergite with a distinct mid-longitudinal ridge (
Fig.
4
in
QUICKE
1994: 229). Hind femur and tibia flattened (
Fig.
3
in QUICKE l.c.;
Fig. 94
). The two
Myosomatoides
species
are keyed after
QUICKE (1994: 230)
Myosomatoides
QUICKE, 1994
2 (3) Wings uniformly brown. (=
myersi
QUICKE, 1994
syn. n.
, =
pennipes
MYERS, 1931
)
M. fasciatus
(
BRULLÉ, 1846
)
3 (2) Wings largely hyaline with apical third of the forewing brown (
Fig.
1
in QUICKE l.c.)
M. pennipes
(
WESTWOOD, 1882
)
4 (1) Second tergite without ridge. Hind femur and tibia not flattened (
Figs 24
,
32
,
40
,
51
,
60
,
69
,
77
,
87
).
5 (6) Hind femur and tibia with short hairs as usually in
Braconinae
(
Fig. 32
). Vein
1r–m
of hind wing weakly bent (
Figs 36
,
45
) to almost straight (
Fig. 44
). See the key to the six species of the genus
Amyosoma
in VAN ACHTERBERG & POLASZEK (1996: 21). The seventh species,
A. cavei
sp. n.
, runs to
A. rufescens
(
QUICKE et INGRAM, 1993
)
with the help of this key, the distinction of these two species is presented at
M. (A.) cavei
sp. n.
Amyosoma
VIERECK, 1913
6 (5) Hind femur and hind tibia with long hairs (
Figs 24
,
51
,
69
,
87
). Vein
1r–m
of hind wing distinctly bent (
Figs 64
,
73
)
Myosoma
BRULLÉ, 1846
7 (10) Fore wing: Second submarginal cell relatively short, i.e.
3–SR
1.5–1.6 times longer than
2–SR
;
SR1
1.5–1.7 times longer than
3–SR
(
Figs 79
,
89
). Temple in dorsal view strongly rounded to receded (
Figs 75, 82
,
85
).
8 (9) First tergite twice longer than broad behind, scutum clearly broadening posteriorly (
Fig. 91
). Head in dorsal view relatively more transverse, 1.7 times as broad as long, temple strongly rounded (
Fig. 85
). Vein
SR1
of fore wing approaching tip of wing (
Fig. 89
). ♀:
6 mm
. –
Brazil
M. rubriventre
BRULLÉ, 1846
9 (8) First tergite five times as long as broad behind, scutum less broadening posteriorly (
Fig. 81
). Head in dorsal view relatively less transverse, 1.6 times as broad as long, temple receded (
Figs 75, 82
). Vein
SR1
of fore wing reaching tip of wing (
Fig. 79
). ♀:
8–8.5 mm
. –
Colombia
M. lagopus
(
KRIECHBAUMER, 1900
)
10 (7) Fore wing: Second submarginal cell relatively long, i.e.
3–SR
1.8–2 times longer than
2–SR
;
SR1
1.1–1.3 times longer than
3–SR
(
Figs 26
,
53
,
62
,
71
). Temple in dorsal view rounded to less rounded (
Figs 22
,
49
,
58
,
67
).
11 (14) Second submarginal cell very long,
3–SR
2.4–2.6 times as long as
2–SR
(
Figs 26
,
53
). Head in dorsal view a bit more transverse, 1.7 times as broad as long (
Figs 22
,
49
)
12 (13) Eye in dorsal view slightly less protruding and 1.5 times as long as temple (
Figs 38–39
). Hind femur less thick, 3.8 times as long as broad medially (
Fig. 40
). Metasoma black, second tergite laterally reddish. Fore wing brown fumous, distally (beyond pterostigma) subhyaline. ♀:
8 mm
. –
Bolivia
M. brullei
SZÉPLIGETI, 1906
13 (12) Eye in dorsal view slightly more protruding and 1.6 times as long as temple (
Fig. 49
). Hind femur thick. 2.9 times as long as broad medially (
Fig. 51
). Metasoma testaceous, tergites 6–8 black. Fore wing brown fumous, basally subhyaline. ♀:
10 mm
. –
Brazil
M. errans
(
SZÉPLIGETI, 1902
)
14 (11) Second submarginal cell long,
3–SR
1.8–1.9 times as long
2–SR
(
Figs 62
,
71
). Head in dorsal view a bit less transverse (
Figs 58
,
67
).
15 (16) Basal lobe of claw deep (
Fig. 70
). Eye in dorsal view 1.7 times longer than temple (
Fig. 67
). Scape in outer-lateral view longer than broad apically (
Fig. 66
). Metasoma reddish. ♀:
8.5 mm
. –
Ecuador
M. hirtipes
BRULLÉ, 1846
16 (15) Basal lobe of claw less deep (
Fig 61
). Eye in dorsal view 1.4 times longer than temple (
Fig. 58
). Scape in outer-lateral view cubic-form, i.e. as long as broad apically (cf.
Fig. 48
). Metasoma testaceous, apically black. ♀:
9 mm
. –
Brazil
,
Belize
M. fuscipenne
BRULLÉ, 1846
Taxonomic remarks – 1.) The genera
Amyosoma
VIERECK, 1913
and
Myosoma
BRULLÉ, 1846
were considered as two valid taxa, however, recently
QUICKE & INGRAM (1993: 317)
and
BELSHAW
et al.
(2001: 423)
formally considered them congeneric. However, the genus
Amyosoma
is a valid genus accepting VAN ACHTERBERG & POLASZEK’ s (1996: 20) point of view on the distinction of the two genera in question.
2.)
Ichneumon mutator
FABRICIUS, 1775
was rearranged by
BRULLÉ (1846: 453)
into the genus
Myosoma
; recently QUICKE & INGRAM designated it as the
type
species of their new genus
Poecilobraconoides
QUICKE et INGRAM, 1993
. Furthermore, the species
Iphiaulax bipartitus
SZÉPLIGETI, 1905
is conspecific with
P. mutator
(FABRICIUS) (
QUICKE 1991: 175
)
distributed in
Australia
.
3.) The sixth species of the genus
Myosoma
,
M. luteum
, described by SZÉPLIGETI in 1913 was placed in the genus
Cratocnema
SZÉPLIGETI, 1914
(
QUICKE 1991: 184
,
PAPP 2000: 155
). Concerning the new combination (
Cratocnema lutea
) QUICKE (l.c.) pointed out: “Very aberrant species of
Cratocnema
...” As a result of the examination of the female
lectotype
of
Myosoma luteum
I confirm QUICKE’ s statement with the supplementary comment that it does not represent the genus
Cratocnema
but, perhaps, it belongs to an undescribed genus.
4.) The present taxonomic status of the
Myosoma
species
by CAMERON see the checklist in Appendix.