Systematic revision of American Glaresidae (Coleoptera: Scarabaeoidea) Author Gordon, Robert D. Author Hanley, Guy A. text Insecta Mundi 2014 2014-01-10 2014 333 1 91 journal article 10.5281/zenodo.4645613 1942-1354 4645613 112A1F0B-1A82-4672-842B-A79A21F251D6 Glaresis howdeni Gordon Glaresis howdeni Gordon 1970: 504 . Description. Male. Length 3.1, width 1.8 mm ; body form elongate, slightly widened from elytral base to apical 1/3 ( Fig. 20A ). Color yellowish brown. Head with clypeal surface and frons densely rugose, impunctate, with large, dense, irregular tubercles, setae short, barely evident; vertex not tuberculate, without trace of basal carina, surface rugose. Clypeal apex weakly emarginate, with small, irregular tubercles, appearing dentate, lateral angles feebly oblique, acute ( Fig. 20B ). Mandible pair symmetrical; mesal tooth strong; lateral prominence strong, pronounced; outer margin rounded. Pronotum with deep, long, transverse fovea in anterior 1/4 extended completely across pronotum, small slightly transverse fovea on each side of middle in anterior 1/2, wide, deep fovea on each side medially near lateral margin, deep central furrow extended from base to transverse furrow; surface densely rugose, with irregular, mostly vertical setae-bearing carinae, setae semidecumbent, about 1/2 length of carina. Anterior and lateral pronotal margins not bordered, all margins except anterior crenulate. Elytra with surface feebly shiny, densely microreticulate; all striae distinctly, weakly carinate, carinae widely interrupted adjacent to space between interval punctures, each carinal segment bearing an apical seta about 2/3 length of segment; intervals with deep round punctures. Metasternum long, dull, finely, densely microreticulate, surface not tuberculate medially, laterally with some elongate, oblique, seta-bearing ridges, median area slightly depressed, with faint trace of carina extended anteriorly from apical keel; metasternal groove weak, distinct, inner margin not ridged, outer margin not ridged ( Fig. 20D ). Lateral protibial teeth unevenly spaced, basal two teeth close together. Mesotibia with 3 short, widely spaced spines in posterolateral emargination, tibia slightly projecting at apex, apically rounded ( Fig. 20D ). Posterior metatrochanteral margin lacking teeth, very slightly serrate, posterosuperior metatrochanteral surface with 1 large tooth ( Fig. 20C ). Metafemoral surface with small, widely scattered, slightly elongate, setaebearing tubercles, microreticulate, slightly shiny throughout; width to length ratio 1.0:1.6, with narrow flange on anterior margin; posterosuperior margin without teeth ( Fig. 20G ). Metatibia broadly triangular, surface entirely microreticulate, with large, bifid, posteromedian lateral projection, row of small teeth anterior to lateral projection, medially with irregular row of coarse tubercles extended from base nearly to apex, inner margin smooth, pubescent ( Fig. 20F ). Apical margin of 5th ventrite broadly, weakly emarginate. Genitalia long, basal piece sligthly longer than parameres, proximal end curved; median lobe longer than parameres, gradually narrowed from near base to acute apex, as wide at middle as a paramere, slightly curved upward in apical 1/3, apex acute in ventral view; paramere straight in lateral view, apex bluntly rounded ( Fig. 20E ). Female . Apex of 5th abdominal ventrite same as in male. Variation . Some specimens have a second, extremely small, barely visible trochanteral tooth. Type locality . California , Kern Co., Alta Sierra, 5800'. Type depository . CNIC. Temporal distribution . June. Geographical distribution . UNITED STATES . California : Inyo Co., 12 mi. E. Big Pine, Inyo Mts., 7700'; Kern Co., Alta Sierra, 5800; Mono Co., Benton Sta.; Mono Co., Larkin Lake; L. Big Rock Cr. P. C.; Siskiyou Co., Horn Ranch. ( CASC ) ( USNM ) Remarks . This species is in subgroup A because of the weakly impressed metasternal groove. It is distinguished from other subgroup members by a combination of small, closely spaced tubercles on clypeal apex; large, dense tubercles on clypeus and frons; posteriosuperior metatrochanteral surface with single large tooth; unique male genitalia; and montane type locality. It is very similar to G . california , but the distinctive male genitalia and differing number of posterosuperior metatrochanteral teeth characterize each. See Remarks under G . california . Gordon (1970) stated that G . howdeni had 2 teeth on the posterosuperior surface of the metatrochanter, but reexamination of the type specimen shows that it has only a single tooth.