A New Genus and Species of Dynastinae (Coleoptera: Scarabaeidae) from the Galápagos Islands, Ecuador, other New Species of Cyclocephalini, Pentodontini, and Phileurini from South America, and a Revised Key to the Genera of New World Pentodontini Author Ratcliffe, Brett C. text The Coleopterists Bulletin 2014 2014-12-31 68 4 663 680 http://dx.doi.org/10.1649/0010-065x-68.4.663 journal article 10.1649/0010-065X-68.4.663 5369368 Hiekeianus halophilus Ratcliffe , new species ( Figs. 11–12 ) Type Material. Holotype female with handwritten label: “ PERU : Cusco /Salineras de Maras/ S 13°18′08″ W 72°9′15″/ 5 Jan 2012 , 3000m / M. Jameson and L. Russell //Found dead in/salt catchment” and with my red holotype label. Type deposited at the University of Nebraska State Museum (Lincoln, NE). Holotype . Female. Length 30.0 mm; width 14.5 mm . Color dark reddish brown. Head: Frons with large, dense, deep, confluent punctures. Frontoclypeal region at center with short, transversely arcuate tubercle. Clypeus triangular, apex narrowly rounded, not reflexed, surface transversely rugose. Mandibles exposed, large, evenly rounded. Interocular width equals 4.0 transverse eye diameters. Antenna 10-segmented, club subequal in length to antennomeres 2–7. Pronotum: Surface convex, lacking fovea or tubercles, weakly crazed, anterior fifth half with moderately large, dense, deep punctures. Remainder of surface with moderately large punctures sparse to moderate in density. Base with thick marginal bead. Elytra: Sutural stria a row of deep, elongate punctures. Surface completely, densely punctate, punctures mostly large, glabrous. Pygidium: Disc shiny, with small, sparse, glabrous punctures. Base with narrow, transverse band of, setigerous punctures; punctures small to moderate, and setae long, reddish brown. In lateral view, surface convex. Legs: Protibia quadridentate. Metatibia at apex expanded, truncate and minutely crenulate with 17 stout spinules. Venter: Prosternal process moderately long, laminate, apex subtruncate. Table 1. Morphological comparison of Stenocrates caiporae and Stenocrates haackae .

Character	S. caiporae	S. haackae
Length	25 mm	21 mm
Clypeal apex	Slightly emarginate	Subtruncate
Frons	Smooth	With large, dense punctures
Interocular width	2.2 transverse eye diameters	2.8 transverse eye diameters
Pronotum	Mostly smooth with small, sparse	Small, sparse punctures on disc with large
punctures in posterior angle	dense punctures in posterior angle
Pygidium	Completely, densely punctate	Sparse punctures on apical half
Protibial teeth	Spaced subequally	Basal tooth slightly removed
Prosternal process	Apex broadly rounded	Apex conical

Etymology. The specific epithet is derived from the Greek words halos for salt and philia for fondness. It is used here in reference to the salt catchments in which it was found. This is not to imply that this species actually has a predilection for salty habitats, but that it was simply found dead in such a place. Distribution. Hiekeianus halophilus is known only from the area of the Maras salt pan area in Cusco Department, Peru . Locality Record. PERU : CUSCO : Salineras de Maras ( Fig. 12 ). Temporal Distribution. January (1). Diagnosis. The female of H. halophilus is described as new because it is so different from the females of H. tuberculatus . The differences between H. halophilus and H. tuberculatus are summarized in Table 2 . In size and gestalt , H. halophilus also resembles Barutus hartmanni Ratcliffe from Panama and Costa Rica , but in B . hartmanni the clypeal apex is broader, the mandibles are weakly lobed, the frontoclypeal region has a binodose tubercle, the mentum is lanceolate rather than constricted at the subapex, the last abdominal sternite is subtruncate, and the prosternal process is large, thick, and rounded as opposed to the laminate, subtruncate process in H. halophilus . Fig. 11. Hiekeianus halophilus , holotype. Fig. 12. Salt catchments at Salineras de Maras, Cusco, Peru, type locality for Hiekeianus halophilus . Photograph by M. L. Jameson. Table 2. Morphological comparison of Hiekeianus halophilus and Hiekeianus tuberculatus .

Character	H. halophilus	H. tuberculatus
Length and depth	30 mm and 13 mm	24–26 mm and 11 mm
Frontoclypeal suture	Tubercle short, transverse, arcuate	Tubercle simple, conical
Clypeal apex	Narrowly rounded	Narrowly bidentate
Mentum surface	Punctures sparse, shallow	Punctures dense, deep
Mentum shape	Lyre-shaped and elongated anteriorly	Lyre-shaped but not elongated anteriorly
Pronotum	Basal bead thick, complete	Basal bead slender, incomplete at center
Elytra	Surface glabrous	With sparse, reddish brown setae,
especially near apices
Last abdominal sternite	Weakly emarginate	Subtruncate

Natural History. The single holotype was found dead, and so nothing is known of its life history.