<document id="C3CAF3142E939E14E9E82E79CA8E3240" ID-DOI="10.5852/ejt.2024.935.2555" ID-ISSN="2118-9773" ID-Zenodo-Dep="11400806" ID-ZooBank="36F07FD1-09B8-4057-86F6-5F6960DC799B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1717145828877" checkinUser="plazi" docAuthor="Bucher, Manon, Bouteille, Emilien &amp; Cesne, Maxime Le" docDate="2024" docId="E8713702D101FFB19266B096FE43FCEB" docLanguage="en" docName="EJT.2024.935.166-202.pdf" docOrigin="European Journal of Taxonomy 935" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2555/11547" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pogonorhinella Schmidt 1910" docType="treatment" docVersion="2" lastPageNumber="185" masterDocId="14484F7AD110FFA2905EB56AFFFDFFE9" masterDocTitle="Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar" masterLastPageNumber="202" masterPageNumber="166" pageNumber="183" updateTime="1717146405090" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FAFAE1E928730790B9DF393F36DB97FE">Malagasy spittlebugs (Hemiptera: Cercopidae), a taxonomic review of genera from Madagascar</mods:title>
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<mods:namePart id="D4CF7567DB4372C28D8E7CCB4F463324">Bucher, Manon</mods:namePart>
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<mods:affiliation id="0C72EB41930AF8EED1CAC5E0EBC81A04">Institut de Systématique, Evolution, Biodiversité (ISYEB), MNHN-CNRS-Sorbonne Université- EPHE-Université des Antilles, Muséum national d’Histoire naturelle, CP 50, 45 rue Buffon, 75005 Paris, France.</mods:affiliation>
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<mods:namePart id="214F445F61A303AF919DDD94BFEBACA9">Bouteille, Emilien</mods:namePart>
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<mods:affiliation id="0A953DBDFB53061110FFE982CDF4B590">Mécanismes adaptatifs et évolution (MECADEV), Muséum national d’Histoire naturelle, CNRS, 57 rue Cuvier, CP 50, 75005 Paris, France.</mods:affiliation>
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<mods:namePart id="63791D0D39ED0C1F9F5F1447F2D4FA66">Cesne, Maxime Le</mods:namePart>
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<mods:affiliation id="0D81393C3E2152693C642B5C689D7611">Institut de Systématique, Evolution, Biodiversité (ISYEB), MNHN-CNRS-Sorbonne Université- EPHE-Université des Antilles, Muséum national d’Histoire naturelle, CP 50, 45 rue Buffon, 75005 Paris, France.</mods:affiliation>
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<paragraph id="60678614D101FFB39266B096FC01F9FF" blockId="17.[568,1020,1531,1593]" box="[568,1020,1531,1558]" pageId="17" pageNumber="183">
<heading id="3B2F3178D101FFB39266B096FC01F9FF" box="[568,1020,1531,1558]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="183" reason="2">
Genus 
<taxonomicName id="A7D8FD97D101FFB392D4B091FC01F9FF" ID-CoL="6TP2" authority="Schmidt, 1910" authorityName="Schmidt" authorityYear="1910" box="[650,1020,1531,1558]" class="Insecta" family="Cercopidae" genus="Pogonorhinella" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="183" phylum="Arthropoda" rank="genus">
<emphasis id="52AC5A06D101FFB392D4B091FCB4F9FF" bold="true" box="[650,841,1531,1558]" italics="true" pageId="17" pageNumber="183">Pogonorhinella</emphasis>
<bibRefCitation id="0449FBE5D101FFB3930FB091FC01F9FF" author="Schmidt E." box="[849,1020,1531,1558]" pageId="17" pageNumber="183" pagination="324 - 340" refId="ref15648" refString="Schmidt E. 1910. Zur Kenntnis indoaustralischer und sudamerikanischer Cercopiden. (Hemiptera- Homoptera). Entomologische Zeitung. Herausgegeben von dem entomologischen Vereine zu Stettin. 71: 324 - 340." type="journal article" year="1910">Schmidt, 1910</bibRefCitation>
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<paragraph id="60678614D101FFB392A9B375FCC1F9D0" blockId="17.[568,1020,1531,1593]" box="[759,828,1567,1593]" pageId="17" pageNumber="183">
<figureCitation id="F8E39A91D101FFB392A9B375FCC1F9D0" box="[759,828,1567,1593]" captionStart="Fig" captionStartId="18.[189,231,1892,1918]" captionTargetBox="[282,1315,290,1824]" captionTargetId="figure-70@18.[247,1339,249,1847]" captionTargetPageId="18" captionText="Fig. 8. Pogonorhinella madagascariensis Schmidt, 1910, ♂ (MNHN(EH) 25271). A. Dorsal view. B. Lateral view. C. Frontal view. D. Labels. E. Male terminalia in lateral view, pygofer, anal tube, aedeagus, left paramere and left subgenital plate. F. Aedeagus. G. Left paramere. H. Distribution map. Scale bars: A–C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11400842" httpUri="https://zenodo.org/record/11400842/files/figure.png" pageId="17" pageNumber="183">Fig. 8</figureCitation>
</paragraph>
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<treatmentCitation id="E179A005D101FFB390E3B30AFD94F993" author="Schmidt E." box="[189,617,1632,1658]" page="334" pageId="17" pageNumber="183" year="1910">
<taxonomicName id="A7D8FD97D101FFB390E3B30AFD94F993" ID-CoL="6TP2" authority="Schmidt, 1910: 334" authorityName="Schmidt" authorityPageNumber="334" authorityYear="1910" box="[189,617,1632,1658]" class="Insecta" family="Cercopidae" genus="Pogonorhinella" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="183" phylum="Arthropoda" rank="genus">
<emphasis id="52AC5A06D101FFB390E3B30AFE8AF993" box="[189,375,1632,1658]" italics="true" pageId="17" pageNumber="183">Pogonorhinella</emphasis>
<bibRefCitation id="0449FBE5D101FFB39120B30AFD94F993" author="Schmidt E." box="[382,617,1632,1658]" pageId="17" pageNumber="183" pagination="324 - 340" refId="ref15648" refString="Schmidt E. 1910. Zur Kenntnis indoaustralischer und sudamerikanischer Cercopiden. (Hemiptera- Homoptera). Entomologische Zeitung. Herausgegeben von dem entomologischen Vereine zu Stettin. 71: 324 - 340." type="journal article" year="1910">Schmidt, 1910: 334</bibRefCitation>
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.
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<paragraph id="60678614D101FFB390E3B3C8FEA5F952" blockId="17.[189,344,1697,1724]" box="[189,344,1697,1724]" pageId="17" pageNumber="183">
<heading id="3B2F3178D101FFB390E3B3C8FEA5F952" bold="true" box="[189,344,1697,1724]" fontSize="11" level="3" pageId="17" pageNumber="183" reason="3">
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<typeStatus id="BF6338B6D101FFB390E3B3C8FF07F955" box="[189,250,1698,1724]" pageId="17" pageNumber="183">Type</typeStatus>
species
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</heading>
</paragraph>
<paragraph id="60678614D101FFB390E3B3BAFC4DF902" blockId="17.[189,944,1744,1771]" box="[189,944,1744,1771]" pageId="17" pageNumber="183">
<taxonomicName id="A7D8FD97D101FFB390E3B3BAFCFAF903" ID-CoL="4KVYP" authority="Schmidt, 1910" authorityName="Schmidt" authorityYear="1910" box="[189,775,1744,1770]" class="Insecta" family="Cercopidae" genus="Pogonorhinella" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="183" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="52AC5A06D101FFB390E3B3BAFDA9F903" box="[189,596,1744,1770]" italics="true" pageId="17" pageNumber="183">Pogonorhinella madagascariensis</emphasis>
<bibRefCitation id="0449FBE5D101FFB39202B3BAFCFAF903" author="Schmidt E." box="[604,775,1744,1770]" pageId="17" pageNumber="183" pagination="324 - 340" refId="ref15648" refString="Schmidt E. 1910. Zur Kenntnis indoaustralischer und sudamerikanischer Cercopiden. (Hemiptera- Homoptera). Entomologische Zeitung. Herausgegeben von dem entomologischen Vereine zu Stettin. 71: 324 - 340." type="journal article" year="1910">Schmidt, 1910</bibRefCitation>
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by monotypy.
</paragraph>
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<heading id="3B2F3178D101FFB390E3B27BFE3BF8C2" bold="true" box="[189,454,1809,1835]" fontSize="11" level="3" pageId="17" pageNumber="183" reason="3">
<emphasis id="52AC5A06D101FFB390E3B27BFE3BF8C2" bold="true" box="[189,454,1809,1835]" pageId="17" pageNumber="183">Synthetic description</emphasis>
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</paragraph>
<paragraph id="60678614D101FFB390E3B22AFCC0F849" blockId="17.[189,1399,1856,1952]" pageId="17" pageNumber="183">
<taxonomicName id="A7D8FD97D101FFB390E3B22AFE8AF8B3" ID-CoL="6TP2" authorityName="Schmidt" authorityYear="1910" box="[189,375,1856,1882]" class="Insecta" family="Cercopidae" genus="Pogonorhinella" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="183" phylum="Arthropoda" rank="genus">
<emphasis id="52AC5A06D101FFB390E3B22AFE8AF8B3" box="[189,375,1856,1882]" italics="true" pageId="17" pageNumber="183">Pogonorhinella</emphasis>
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is a monospecific endemic genus of 
<collectingCountry id="18CFC684D101FFB39374B22AFC45F8B3" box="[810,952,1856,1882]" name="Madagascar" pageId="17" pageNumber="183">Madagascar</collectingCountry>
which possesses a peculiar shape due to its small head and the narrow anterior margin of the pronotum. A synthesis based on 
<bibRefCitation id="0449FBE5D101FFB39495B209FA8AF894" author="Schmidt E." box="[1227,1399,1891,1917]" pageId="17" pageNumber="183" pagination="324 - 340" refId="ref15648" refString="Schmidt E. 1910. Zur Kenntnis indoaustralischer und sudamerikanischer Cercopiden. (Hemiptera- Homoptera). Entomologische Zeitung. Herausgegeben von dem entomologischen Vereine zu Stettin. 71: 324 - 340." type="journal article" year="1910">Schmidt, 1910</bibRefCitation>
description and material examined is presented below:
</paragraph>
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<paragraph id="60678614D102FFB090E3B20EFE1EF800" blockId="18.[189,1399,1892,2025]" pageId="18" pageNumber="184">
<emphasis id="52AC5A06D102FFB090E3B20EFEF2F896" bold="true" box="[189,271,1892,1919]" pageId="18" pageNumber="184">Fig. 8.</emphasis>
<taxonomicName id="A7D8FD97D102FFB09143B20FFC8DF896" authority="Schmidt, 1910" authorityName="Schmidt" authorityYear="1910" box="[285,880,1893,1919]" class="Insecta" family="Cercopidae" genus="Pogonorhinella" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="184" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="52AC5A06D102FFB09143B20FFD4CF896" box="[285,689,1893,1919]" italics="true" pageId="18" pageNumber="184">Pogonorhinella madagascariensis</emphasis>
<bibRefCitation id="0449FBE5D102FFB092E0B20FFC8DF896" author="Schmidt E." box="[702,880,1893,1919]" pageId="18" pageNumber="184" pagination="324 - 340" refId="ref15648" refString="Schmidt E. 1910. Zur Kenntnis indoaustralischer und sudamerikanischer Cercopiden. (Hemiptera- Homoptera). Entomologische Zeitung. Herausgegeben von dem entomologischen Vereine zu Stettin. 71: 324 - 340." type="journal article" year="1910">Schmidt, 1910</bibRefCitation>
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, ♂ (MNHN(EH) 25271). 
<emphasis id="52AC5A06D102FFB094EBB20EFB37F897" bold="true" box="[1205,1226,1892,1918]" pageId="18" pageNumber="184">A</emphasis>
. Dorsal view. 
<emphasis id="52AC5A06D102FFB090E3B2E2FF2DF84B" bold="true" box="[189,208,1928,1954]" pageId="18" pageNumber="184">B</emphasis>
. Lateral view. 
<emphasis id="52AC5A06D102FFB091D5B2E2FE5DF84B" bold="true" box="[395,416,1928,1954]" pageId="18" pageNumber="184">C</emphasis>
. Frontal view. 
<emphasis id="52AC5A06D102FFB09202B2E2FD8CF84B" bold="true" box="[604,625,1928,1954]" pageId="18" pageNumber="184">D</emphasis>
. Labels. 
<emphasis id="52AC5A06D102FFB092BBB2E2FD05F84B" bold="true" box="[741,760,1928,1954]" pageId="18" pageNumber="184">E</emphasis>
. Male terminalia in lateral view, pygofer, anal tube, aedeagus, left paramere and left subgenital plate. 
<emphasis id="52AC5A06D102FFB09358B2C1FCEBF82C" bold="true" box="[774,790,1963,1989]" pageId="18" pageNumber="184">F</emphasis>
. Aedeagus. 
<emphasis id="52AC5A06D102FFB093F9B2C1FC43F82C" bold="true" box="[935,958,1963,1989]" pageId="18" pageNumber="184">G</emphasis>
. Left paramere. 
<emphasis id="52AC5A06D102FFB094DDB2C1FB65F82C" bold="true" box="[1155,1176,1963,1989]" pageId="18" pageNumber="184">H</emphasis>
. Distribution map. Scale bars: A–C = 1 mm.
</paragraph>
</caption>
<paragraph id="60678614D103FFB190E3B463FD7AFE80" blockId="19.[189,1399,265,362]" pageId="19" pageNumber="185">HEAD. Small with short vertex. Postclypeus narrow with strong carina and striated on sides, producing obtuse angle in lateral view. Rostrum reaching posterior margin of metacoxae. Antennae long, first segment very large, prominent pedicel.</paragraph>
<paragraph id="60678614D103FFB190E3B4FBFEBDFE27" blockId="19.[189,1398,401,462]" pageId="19" pageNumber="185">THORAX. Anterior half of pronotum trapezoidal giving impression of neck. Hindlegs with one spine on metatibiae.</paragraph>
<paragraph id="60678614D103FFB190E3B49CFD8FFDE6" blockId="19.[189,626,501,528]" box="[189,626,501,528]" pageId="19" pageNumber="185">TEGMINA. Tegmina longer than wide.</paragraph>
<paragraph id="60678614D103FFB190E3B75DFAB2FDB8" blockId="19.[189,1359,567,593]" box="[189,1359,567,593]" pageId="19" pageNumber="185">MALE GENITALIA. Male genitalia without sterno-lateral plates. Aedeagus longer than pygofer height.</paragraph>
<paragraph id="60678614D103FFB190E3B712FE36FD7B" blockId="19.[189,459,632,658]" box="[189,459,632,658]" pageId="19" pageNumber="185">
<emphasis id="52AC5A06D103FFB190E3B712FEABFD7B" bold="true" box="[189,342,632,658]" pageId="19" pageNumber="185">Distribution</emphasis>
(
<figureCitation id="F8E39A91D103FFB19138B712FE3CFD7B" box="[358,449,632,658]" captionStart="Fig" captionStartId="18.[189,231,1892,1918]" captionTargetBox="[282,1315,290,1824]" captionTargetId="figure-70@18.[247,1339,249,1847]" captionTargetPageId="18" captionText="Fig. 8. Pogonorhinella madagascariensis Schmidt, 1910, ♂ (MNHN(EH) 25271). A. Dorsal view. B. Lateral view. C. Frontal view. D. Labels. E. Male terminalia in lateral view, pygofer, anal tube, aedeagus, left paramere and left subgenital plate. F. Aedeagus. G. Left paramere. H. Distribution map. Scale bars: A–C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11400842" httpUri="https://zenodo.org/record/11400842/files/figure.png" pageId="19" pageNumber="185">Fig. 8H</figureCitation>
)
</paragraph>
<paragraph id="60678614D103FFB190E3B7CDFDD1FD28" blockId="19.[189,556,679,705]" box="[189,556,679,705]" pageId="19" pageNumber="185">
Endemic genus of 
<collectingCountry id="18CFC684D103FFB191C6B7CDFDD5FD28" box="[408,552,679,705]" name="Madagascar" pageId="19" pageNumber="185">Madagascar</collectingCountry>
.
</paragraph>
<paragraph id="60678614D103FFB190E3B782FE43FCEB" blockId="19.[189,446,744,770]" box="[189,446,744,770]" pageId="19" pageNumber="185">Number of species: 1.</paragraph>
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