A new species of Lasiosmylus from the Early Cretaceous, China clarifies its genus-group placement in Ithonidae (Neuroptera) Author Zheng, Bingyu Author Ren, Dong Author Wang, Yongjie text ZooKeys 2016 636 41 50 http://dx.doi.org/10.3897/zookeys.636.10103 journal article http://dx.doi.org/10.3897/zookeys.636.10103 1313-2970-636-41 168C6C3A9099468DA9743747CB6550F7 168C6C3A9099468DA9743747CB6550F7 Lasiosmylus longus sp. n. Figs 3, 4 Material. Holotype, CNU-NEU-LB2015003, a partly preserved specimen. Body barely preserved, but four overlapping, sub-complete wings, partially folded, with visible features. Diagnosis. Humeral veinlet recurrent, with a few branches; numerous markings present on the forewing; a distinct oblique stripe parallel to the outer margin; costal crossveins simple; one basal subcostal crossvein; Sc and R1 separate distally, Sc terminating in costal margin 2/3 length of wing; R1 with numerous anteriorly directed branches; Rs with more than ten branches; MP fork level with origin of MA; CuA pectinately branched, CuP with three distal branches. Description. Body: ca. 16.3 mm long; head hypognathous, retracted into pronotum partly; antenna filiform (ca. 4.0 mm) and incompletely preserved; compound eye large, ocelli absent; pronotum quadrate, numerous long setae concentrated laterally; mesonotum and metanotum stout; abdomen and legs indiscernible. Fore wing: ca. 22.7 mm long, 7.9 mm wide; slender and membranous with numerous fuscous spots; humeral plate discernible (Fig. 3A); veins covered by dense setae, particular along wing margin; trichosors and nygmata undetectable; costal space broad basally (maximum width = 2.1 mm), narrowed distally; recurrent humeral veinlet with several branches; costal crossveins simple and with the occasional distal dichotomous forks, densely arranged distally; Sc and R1 separated distally; one subcostal crossvein close to the origin of Rs; R1 with many pectinately branches distally, entering the anterior margin; Rs branches regularly arranged with about thirteen branches; few crossveins present between branches of Rs; MA simple; MP first fork distant from wing base, close to the MA divergence from Rs; one mp1-mp2 crossvein detected; CuA branched near the middle of wing, with ten pectinate branches; CuP with three simple branches; anal veins partly preserved, 1A with three branches and forked proximally, 2A proximally forked. Hind wing: ca. 18.0 mm long, 7.3 mm wide, partly preserved, venation similar to forewing except costal space narrow; cubitus veins and anal veins not well preserved (Figs 3, 4). Figure 3. Lasiosmylus longus sp. n. (holotype CNU-NEU-LB2015003). A habitus photograph Bhp, humeral plate (left hindwing) C line drawing. Scale bars: 5 mm (A, C), 1 mm (B). Figure 4. Lasiosmylus longus sp. n. (holotype CNU-NEU-LB2015003), line drawings. A right forewing B left hind wing C right hind wing. Scale bars: 5 mm ( A-C ). Etymology . The species name is from the Latin ' longus ', referring to the slender wing of this moth lacewing. Type locality. Huangbanjigou Village, Beipiao City, Liaoning Province, China. Type horizon. Yixian Formation, Barremian-early Aptian (129.7-122.1 Ma), Early Cretaceous. Remarks. Lasiosmylus longus sp. n. can be distinguished from Lasiosmylus newi by the distinct oblique stripe close to the outer margin, multiple Rs branches, and pectinate CuA branches.