Akrokolioplax, a new genus of Southeast Asian labeonine fishes (Teleostei: Cyprinidae).
Author
E Zhang
Author
Maurice Kottelat
text
Zootaxa
2006
1225
21
30
http://www.zoobank.org/urn:lsid:zoobank.org:pub:F1EE8A2C-1A31-4A02-AE61-9F33857A85EC
journal article
z01225p021
[[ Genus
Epalzeorhynchos Bleeker
]]
The Labeoninae (sensu Chen et al., 1984), which is essentially equivalent to the Labeines of Reid (1982) or the tribe Labeone (= Labeonini) of Rainboth (1991, 1996), includes cyprinid fishes widely distributed in the fresh waters of tropical Africa and Asia. Most of these fishes are adapted to fast flowing waters. Members of this subfamily show a high degree of morphological modification in oromandibular structures, which is the basis for recognition of most of its included genera. To date, twenty-seven Labeonine genera have been recognized from eastern and southeastern Asia (Rainboth, 1991; Zhang & Chen, 2004). Among these,
Epalzeorhynchos Bleeker, 1855
(type species:
Barbus kalopterus Bleeker, 1851
) is diagnosed by having a pair of moveable lateral lobes on the snout, which are also called “fleshy rostral papilla” (Roberts, 1989) or “posteriorly free lateral lobes” (Yang & Winterbottom, 1998). Five valid species have so far been identified in
Epalzeorhynchos
:
E. bicolor
,
E. bicornis
,
E. frenatus
,
E. kalopterus
, and
E. munensis
.
Epalzeorhynchos bicornis
was described by Wu et al. (1977), based on a single 140 mm SL specimen from the upper Salween River (= Nu Jiang in Chinese) basin in Liuku County, Yunnan Province, China. Although
E. bicornis
is currently regarded as valid (Chu & Cui, 1989; Chen, 1998; Zhang et al., 2000), its generic position is still unclear. Kottelat (1989) commented that
E. bicornis
does not belong in
Epalzeorhynchos
, but did not elaborate further. Yang and Winterbottom (1998) investigated the relationships among the five currently identified species of
Epalzeorhynchos
. They demonstrated that
E. bicornis
is sister to
Paracrossochilus
when
Crossocheilus
is used as the outgroup, and that it is basal to the clade that includes
Crossocheilus
and other species of
Epalzeorhynchos
, when
Paracrossochilus
is utilized as the outgroup. These findings led Yang and Winterbottom to conclude that
E. bicornis
should not be placed in
Epalzeorhynchos
, but they stopped short of erecting a new genus for its reception.
Kottelat’s (1989) and Yang and Winterbottom’s (1998) observation that
E. bicornis
is not immediately associated with the other species of
Epalzeorhynchos
is corroborated by marked differences we found in position and structure of the moveable lateral lobes on the snout and the morphology of the oromandibular structures. Morphology of the oromandibular structures is diagnostic for many Labeoninae genera (Zhang et al., 2000; Zhang & Chen, 2004), and the differences we observed among species of
Epalzeorhynchos
indicate that
E. bicornis
belongs to a distinct genus, which we describe and name in the present paper.