Revisions and key to the Vernonieae (Compositae) of Thailand
Author
Bunwong, Sukhonthip
Maejo University Phrae Campus, Mae Sai, Rong Kwang, Phrae 54140, Thailand
Author
Chantaranothai, Pranom
Applied Taxonomic Research Center, Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand
Author
Keeley, Sterling C.
Department of Botany, University of Hawaii, Honolulu, HI 96816 USA
text
PhytoKeys
2014
2014-05-13
37
25
101
http://dx.doi.org/10.3897/phytokeys.37.6499
journal article
http://dx.doi.org/10.3897/phytokeys.37.6499
1314-2003-37-25
FFE8FFACFF84FFA95573FFFECD03F742
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Acilepis silhetensis (DC.) H.Rob., Proc. Biol. Soc. Washington 112(1): 227. 1999.
Vernonia bracteata
Wall., Numer. List [Wallich] no. 2921, comp. no. 31,
nom. nud.
Decaneurum silhetense
DC., Prodr. 5: 67. 1836.
Vernonia silhetensis
(DC.) Hand.-Mazz., Symb. Sin. 7: 1084. 1936.
Type.
India, Silhet;
Wallich
2921 (holotype: E!).
Fig. 5H
.
Description
.
Perennial herbs, 1-3 m tall. Stems erect, inconspicuously ribbed, pilose. Leaves 7-12 by 2-4 cm, lanceolate or oblanceolate, margin serrate, apex acute or acuminate, base attenuate, subcoriaceous; both surfaces scabrous with whip-shaped hairs and capitate glands; lateral veins 5-10-paired; petioles up to 1 cm long. Capitulescences terminal, paniculate or solitary. Capitula campanulate, 15-20 mm long, pedunculate. Receptacle flat, 6-10 mm in diam., hairy. Involucres in 6-7 series, 11-18 mm long, 10-15 mm in diam., herbaceous, campanulate. Phyllaries dark purple or green with purple apex, margin piliferous, outer surface arachnoid lacking glands; the outer and the middle ones ovate, apex acuminate, upper half strongly reflexed; the inner ones ovate-lanceolate to oblong, apex acuminate or aristate. Florets 50-75; corollas funnelform, purple, glandular; corolla tubes 8-12 mm long; corolla lobes 3.5-5 mm long. Anthers 3.5-4.5 mm long, apical appendage acute, base obtuse. Styles purple, 10-12 mm long, branches 3.5-4.5 mm long. Achenes subterete, 4-5 mm long, 10-ribbed, glandular. Pappus bristles, the inner ones 6.5-8 mm long.
Distribution.
Thailand: Mae Hong Son, Chiang Mai, Lamphun, Lampang, Phetchabun, Loei, Khon Kaen. China, India, Bhutan, Myanmar, Laos, Myanmar.
Specimens examined.
Thailand, Chiang Mai, Queen Sirikit botanic garden,
18°53.89'N
,
98°51.61'E
, 10 Dec 2007,
S. Bunwong
364 (KKU, US); Mae Hong Son, Pai, 20 Oct 1979,
T. Shimizu, H. Toyokuni, H. Koyama, T. Yahama & T. Santisuk
T-20113 (BKF, L); Chiang Mai, Doi Chiangdao, Nov 25 1963,
Adisai
714 (BK); Doi Chiangdao, 24 Jul 1998,
K. Buchuai
279 (AAU, BKF); Doi Chiangdao, 16 Dec 1983,
N. Fukuoka & M. Ito
T-35208 (BKF); Doi Chiangdao,3 Dec 1984,
H. Koyama
T-39781 (BKF, L); Doi Chiangdao, 5 Aug 1988,
H. Koyama
T- 61104 (BKF); Doi Chiangdao,29 Nov 1984,
H. Koyama, T. Yahara, H. Nagamasu, W. Nanakorn & N. Nantasan
T-39710 (AAU, BKF, L); Doi Chiangdao, 30 Nov 1984,
H. Koyama, T. Yahara, H. Nagamasu, W. Nanakorn & N. Nantasan
T- 39736 (BKF, L); Doi Chiangdao, 4 Nov 1995,
J.F. Maxwell
95-1065 (CMU, BKF); Doi Chiangdao, 13 Oct 1931,
Put
311 (BK, BM, K); Chiang Mai, Doi Sutep, 6 Oct 1982,
Konta Th. Wongprasert & B. Sangkhachand
29741 (BKF); Doi Sutep,28 Nov 1984,
H. Koyama, S. Mitsuta, T. Yahara & H. Nagamasu
T-39671 (BKF, L); Doi Inthanon, 6 Dec 1984,
S. Mitsuta, T. Yamada & H. Nagamazu
T-46454 (BKF); Doi Inthanon, 1 Oct 1971,
G. Murata, K. Iwatsuki, C. Phengklai & C. Charoenpol
T-15497 (BKF); Doi Inthanon, 1 Oct 1971,
G. Murata, K. Iwatsuki, C. Phengklai & C. Charoenpol
T-15498 (BKF, P); Mae Rim, 21 Dec, 1985,
Y. Paisooksantivatana
1645b-85 (BK); Mae Rim, 26 Dec 1987,
R. Pooma
43 (BKF); Lampang, Jae son national park, 2 Dec 1995,
J.F. Maxwell
95-1227 (BKF, L); Doi Khun Tan, 2 Jan 1985,
H. Koyama & C. Phengklai
T-39197 (BKF, L); Phetchabun, Namnao national park, 18 Nov 1973,
J.F. Maxwell
73-636 (AAU, BK, BKF); Loei, Phu Kradung, 24 Dec 1991,
Dee
195 (E); Phu Kradung, 21 Oct 1989,
Din
125 (BKF); Phu Kradung, 19 Dec 1982,
H. Koyama, H. Terao & Th. Wongprasert
T-31330 (C, BKF); Phu Kradung, 5 Sep 1988,
R. Pooma
62 (BKF); Phu Kradung, 21 Oct 1967,
Prayad
1076 (BK); Khon Kaen, route to Nam Nao national park, 18 Nov 1979,
T. Shimizu, H. Toyokuni, H. Koyama, T. Yahama & T. Santisuk
T-22531 (BKF, L).
Diagnostic
characters.
This species is distinguished by its strongly reflexed phyllaries and glandular achenes lacking trichomes.
Ecology.
Dipterocarp or pine-oak forest, alt. 700-1350 m; flowering August to January.
Vernacular names.
Phak Phet Kao Kum (ผักเผ็ดข้าวกํ่า), Ya Klung (หญ้าคลัง), Ya Hang Nok Kiew (หญ้าหางนกเขียว), Hudsakuen (หัสคึน).