A taxonomic revision of the genus Noronhia Stadtm. ex Thouars (Oleaceae) in Madagascar and the Comoro Islands
Author
Hong-Wa, Cynthia
Missouri Botanical Garden P. O. Box 299 St. Louis MO 63166 - 0299 U. S. A & Claude E. Phillips Herbarium Delaware State University 1200 N. DuPont Hwy, Dover DE 19901 - 2277 U. S. A.
chwa@desu.edu
text
Boissiera
2016
2016-10-21
70
1
292
journal article
22287
10.5281/zenodo.7599432
f2ceb54b-e71d-4433-9e67-b75b62f8a902
978-2-8277-0086-8
0373-2975
7599432
48.
Noronhia luteola
H. Perrier
in Mém. Inst. Sci.
Madagascar
, Sér. B, Biol. Vég. 2:
298. 1949 (
Fig. 33A
).
Lectotypus
(designated here):
MADAGASCAR
. Prov.
Toliara
:
Ampandrandava
,
entre Bekily et Tsivory
, [
24°05’S
45°52’E
],
700 m
,
I.1944
,
Seyrig 642b
(
P
[
P0071219
]!, isolecto-:
P
[
P03558932
,
P03558933
]!)
.
Syntypi
:
MADAGASCAR
.
Prov.
Fianarantsoa
:
Vallées de l’Isalo
,
Fanjahira
, [
23°29’S
44°44’E
],
300 m
,
9.X.1924
,
Humbert 2760
(
K
[
K000233187
],
P
[
P00701221
,
P00701273
]!)
.
Prov.
Toliara
: Mont Amboahangy près d’Esira, [
24°15’S
46°39’E
],
1000-1150 m
,
25.XI.1928
,
Humbert 6825
(
G
[00188779] image seen,
P
[
P00701222
]!)
;
Mont Vohipolaka au
N
de Betroka
(Centre-Sud), [
23°08’S
46°05’E
],
1100-1200 m
,
1.XI.1933
,
Humbert 11641
(
P
[
P00701220
,
P03558930
,
P03558931
]!)
;
Mont Vohitrosy, Anadabolava, [
24°15’S
46°43’E
],
800 m
,
XII.1933
,
Humbert 12656
(
P
[
P03558938
]!)
;
Vallée de la Sakamalio
, bassin
du Mandrare
[
24°32’S
46°41’E
],
900-1100 m
,
XII.1933
,
Humbert 13370
(
P
[
P03558934
]!)
;
Ampandrandava,
entre Bekily et Tsivory
, [
24°05’S
45°52’E
],
700 m
,
IV.1943
,
Seyrig 642
(
P
[
P03558935
]!)
.
Description
Trees
to
13 m
tall, trunk to
25 cm
diameter; young twigs cylindrical,
0.7-1.6 mm
diameter, glabrous; bark medium gray, rugose.
Leaves
opposite, deciduous; bud scales persistent; blades medium green glossy above, lighter below, oblong to obovate, 3-12.5
3
1-3 cm
, coriaceous, glabrous, domatia absent, base attenuate, margin slightly revolute, apex rounded to acute or slightly acuminate, the acumen
1-9 mm
long, midrib slightly sunken above, slightly raised below, secondary veins barely visible, 7-16 per side,
4-13 mm
apart, looping
0.8-3.5 mm
from the margin; petiole yellow to reddish, 3-10
3
0.7-1.7 mm
, not woody, glabrous.
Thyrses
geminate, pauciflorous, compact to somewhat diffuse; peduncle
2 mm
long, glabrescent; pedicel
1.5-18 mm
long, glabrescent; calyx sparsely pubescent outside, glabrous inside, lobes triangular, 0.7-1.3
3
1-1.5 mm
; corolla white, subrotate,
3-4 mm
long, glabrous on both sides, the tube
1-3 mm
long, lobes widely ovate, apex rounded; corona present,
1-1.7 mm
long, undivided; stamens
1.5-2.6 mm
long, anthers transversely oblong,
1.1-1.8 mm
long; pistil
1.5-2.2 mm
long, stigma capitate.
Fruiting
pedicel 3-7
3
0.9-1.1 mm
; young fruits green, brown when mature, ovoid, 11-19
3
8-12.5 mm
, surface smooth, apex flat to rostellate, the rostellum flattened, with the persistent style; dry pericarp
0.6-1.2 mm
thick; endocarp woody; seed 7-13
3
5.5-8.5 mm
.
Distribution, ecology and phenology
Noronhia luteola
occurs in littoral and low- to high-elevation dry forests and thickets in the south, from Fanjahira to Fort-Dauphin (
Fig. 27
). It produces flowers and fruits from October to June.
164
Boissiera 70
Conservation status
The assessment included 23 collections representing 13 localities and resulted in an EOO of
21,499 km
2
, an AOO of
44 km
2
, and eight subpopulations representing seven locations, of which two occur within the newly established protected areas of Mandena and Petriky. Some of the subpopulations occur in an area that is frequently burned for grazing while others are currently affected by industrial mining in southern
Madagascar
. Because of observed and projected continuing decline in habitat quality as well as habitat loss, coupled with an actual AOO that is probably less than
2,000 km
2
,
N. luteola
is assigned a preliminary status of “Vulnerable” [VU B2ab(ii,iii,iv,v)], although the EOO is not expected to decline in the near future.
Notes
Noronhia luteola
can be recognized by its coriaceous, oblong to obovate leaf blades, short-peduncles, and subrotate, white flowers. The following
syntypes
were cited in the protologue:
Humbert 2760
,
6825
,
11641
,
12656
and
13370
,
Seyrig 642
and
642b
. Of these,
Seyrig 642b
is the only one with flowers and several duplicates, and one of the sheets at P has therefore been chosen as the
lectotype
.
Additional specimens examined
MADAGASCAR
.
Prov.
Toliara
:
Forêt de Petriky
,
25°05’S
46°52’E
,
0-10 m
,
5.XII.1989
,
Dumetz
&
McPershon
1121
(
MO
,
TAN
)
;
Ampasy Nampoana
, forêt de
Mandena
,
24°57’10’’S
47°00’10’’E
,
9 m
,
12.
II
.2009
,
Hong-Wa 594
(
MO
,
P
,
TAN
)
;
Mandena forest
,
24°57’S
47°00’E
,
25 m
,
25.X.1989
,
McPherson
et al. 14230
(
MO
,
P
,
TAN
,
TEF
)
;
Manambaro
,
Ambovo
, forêt de
Petriky
,
25°03’03’’S
46°53’44’’E
,
6 m
,
24.
VI
.2004
,
Rabenantoandro
et al. 1606
(
G
,
MO
,
P
,
TEF
)
;
Manafiafy
,
24°46’S
47°12’E
,
12.XI.1990
,
Rabevohitra
2434
(
K
,
MO
,
P
,
TAN
,
TEF
)
;
Mahatalaky
,
Belavenoka
,
24°50’S
47°04’E
,
15.XI.2007
,
Ramison
&
Rabevohitra
400
(
G
,
MO
,
P
,
TAN
)
.