Systematic revision of the pantropical whip spider family Charinidae Quintero, 1986 (Arachnida, Amblypygi) Author Miranda, Gustavo Silva de 81150D94-592A-4CE5-8E88-E60F557A4341 Center for Macroecology, Evolution and Climate, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark. & Entomology Department, National Museum of Natural History, Smithsonian Institution, Washington DC, USA. smiranda.gustavo@gmail.com Author Giupponi, Alessandro P. L. 434112AC-B212-43E8-A5D9-2F5D5619AFC4 Laboratório de Referência Nacional em Vetores das Riquetsioses, LIRN-IOC-FIOCRUZ, Rio de Janeiro, RJ, Brazil. agiupponi@gmail.com Author Prendini, Lorenzo C2D080D0-75DB-4DA1-A101-AB4DCF50FF0A Arachnology Lab, Division of Invertebrate Zoology, American Museum of Natural History, New York, NY, USA. lorenzo@amnh.org Author Scharff, Nikolaj F84D2235-66D2-460C-820D-80024068759D Center for Macroecology, Evolution and Climate, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark. & Entomology Department, National Museum of Natural History, Smithsonian Institution, Washington DC, USA. & Zoology Section, Natural History Museum of Denmark, University of Copenhagen, Denmark. nscharff@snm.ku.dk text European Journal of Taxonomy 2021 2021-09-24 772 1 409 http://dx.doi.org/10.5852/ejt.2021.772.1505 journal article 4042 10.5852/ejt.2021.772.1505 b65bb5a9-bbe7-49a4-af44-4b4c03121288 2118-9773 5536410 9B82A32F-0A07-47E3-8684-FED7C8EBF1E9 Charinus tomasmicheli Armas, 2006 Fig. 12 ; Table 1 Charinus tomasmicheli Armas, 2006a: 168–169 , figs 1–4. Charinus tomasmicheli – Armas et al. 2009: 136 ; 2013a: 16 . — Teruel & Questel 2015: 47 . —Rodríguez- Cabrera & Teruel 2016: 1–7 , figs 1–7. — Miranda et al. 2016b: 555 , 557. Diagnosis Based on the description of a male byArmas (2006a), this species may be separated from other Caribbean and Central American Charinus by means of the following combination of characters: almost uniform light yellowish-brown color in life; median eyes and ocular tubercle absent; lateral eyes pigmented; tritosternum short and narrow with typical setation; bifid tooth of basal cheliceral segment with cusps equal in size; cheliceral claw with four teeth; pedipalp femur with three dorsal spines and three ventral spines; pedipalp femur with two setiferous tubercles proximal to first dorsal spine; pedipalp patella with three dorsal spines and two ventral spines; pedipalp patella with large setiferous tubercle between spine I and distal margin; pedipalp tarsus with two dorsal spines; tibia of leg I with 21 articles, tarsus I with 37 articles; first tarsal article same length as subsequent articles; leg IV basitibia with three pseudoarticles; trichobothrium bt situated in proximal third of distitibia; trichobothrium bc situated closer to sbf than to bf . Only adult males are known (all females are immature). Charinus tomasmicheli occurs in the same mountain range as C. centralis , in which 23 tibial articles and 41 tarsal articles are present on leg I. Compared to other Cuban species, C. tomasmicheli possess the same number of articles on the tibia of leg I and the median eyes and ocular tubercle are absent as in C. cubensis and C. wanlessi . However, the leg I tarsus consists of 33 articles in C. wanlessi , whereas 37 articles are counted in C. tomasmicheli ; the first article of the leg I tarsus is around twice the length of the subsequent articles in C. cubensis , whereas these articles are about the same length in C. tomasmicheli . Charinus tomasmicheli is very similar to C. acosta , in which the median eyes are present and the cheliceral claw bears five teeth instead of four. Etymology Patronym honoring Tomás Michel Rodriguéz, one of the collectors of the type series ( Armas 2006a ). Type material Holotype CUBA • ♂ ; Villa Clara Province , Manicaragua Municipality , Los Manantiales , Guamuhaya Massif , Los Manantiales-Tito cave ; 21°59′ N , 80°01′ W ; 470 m a.s.l. ; 11 Mar. 2005 ; T.M. Rodríguez and R. Chaviano leg.; on wall in dark area near stream ; IES [not examined]. Paratypes CUBA • 1 ♂ ; same collection data as for holotype but 15 Oct. 2004 ; on an accumulation of sediments ; IES [not examined] • 1 subad. ♀ ; same collection data as for holotype but 16 Oct. 2004 ; in small cavity in wall, twilight zone ; IES [not examined] • 3 subad. ♀♀ , 1 ♂ , 1 juv. ♀ ; same locality as for holotype; 17 Feb. 2006 ; T.M. Rodríguez leg.; twilight zone ; IES [not examined] . Measurements See Table 1 . Distribution Distributed in caves and forests in the Cuban provinces of Cienfuegos , Sancti Spíritus , and Villa Clara . Natural history The type series was collected in the Los Manantiales-Tito Cave, in the twilight and dark zones. Specimens were caught on the ground and the walls. The proportionally longer legs in relation to the body suggest this species is troglobitic ( Armas 2006a ). However, Armas (2013a) and Rodríguez-Cabrera & Teruel (2016) report finding C. tomasmicheli in a semi-dark vestibular zone and in epigean areas. Charinus tomasmicheli is sympatric with Paraphrynus viridiceps at most localities; at Caletón de Don Bruno, C. tomasmicheli is sympatric with Phrynus marginemaculatus and, at La Tatagua and Palma Sola, with Phrynus decoratus Teruel & Armas, 2005 ( Armas 2006a ; Rodríguez-Cabrera & Teruel 2016 ).