Litthabitellidae: a new family of the Truncatelloidea (Mollusca: Caenogastropoda) Author Falniowski, Andrzej Author Jaszczyńska, Aleksandra Author Osikowski, Artur Author Hofman, Sebastian text Journal of Natural History 2023 2023-03-13 57 5 - 8 299 329 http://dx.doi.org/10.1080/00222933.2023.2168573 journal article 10.1080/00222933.2023.2168573 83f7b49d-c03b-4658-883f-abbd0c538834 1464-5262 7778206 Litthabitella cetinensis Osikowski et Jaszczyńska sp. nov. ( Figures 3 (b), 4 (t–v), 5 (c,f,i), 9(f)) urn:lsid:zoobank.org:act: 10089647-36C6-4773-8E7F-AA0D5BB02833 Diagnosis Shell small, ovate conical, distinguished from L. chilodia and L. aquaadcrucem by smaller and less thick-walled shell and penis with less broad tip and longer and broader filament. COI sequences: ON661294–ON661297; H3 sequences: KY215954–KY215958; 18S sequences: ON668020–ON668023. COI MDCs: 6, A; 21, G; 39, A; 51, C; 84, G; 90, C; 111, A; 132, G; 135, A; 144, T; 153, C; 171, G; 174, C; 183, G; 219, G; 276, C; 309, C; 369, G; 399, T; 402, C; 436, C; 438, A. Etymology The specific name cetinensis reflects the geographic localisation of the type locality within the valley of Cetina River. Figure 1. Localities of origin of the sequenced specimens. Numbers correspond to Table 1. Colours correspond to genetic lineages in the phylogram (Figure 10). Insert: palaeogeographic map of the Late Tortonian (8–7 Mya), indicating position of continental, shallow and deep basins (modified from Biltekin 2010 ); the red arrow points to the study area. Type material Holotype . CROATIA : ethanol-fixed specimen ( Figure 3 (b)); Kostanje , a medium-sized spring at the right bank of the Cetina River ; 43.4411°N , 16.8228°E , 133 m asl ; our locality 1 ( Figure 2 (a)); June 2011 ; leg. Magdalena Szarowska , Andrzej Falniowski ; MNHW-1440. Paratypes . CROATIA • 10 specs (ethanol-fixed); same collection data as for holotype, DMJU 2212 . Description Shell ( Figure 3 (b)) ovate conical up to 2.02 mm tall, growing regularly. Holotype measurements: see Table 4 . Teleoconch whorls slightly convex, evenly rounded. Aperture narrow, ovate, weakly angled adapically, peristome continuous, in its upper part fitting tight to body whorl. Parietal lip complete, adnate, umbilicus nearly completely covered by lip. Outer lip fluted, orthocline. Shell white and glossy with very slightly marked growth lines, moderately thick-walled and translucent, periostracum white. Initial part of the protoconch narrow, on the protoconch surface only delicate irregularities ( Figure 5 (c)). Measurements of holotype and sequenced and illustrated shells: see Table 4 . Shell variability slight ( Figure 4 (t–v)). Morphology and anatomy of soft parts. The head with eyes and no pigment; mantle pigmented less intensively than in the other Litthabitella . Ctenidium with 14 short lamellae, the osphradium broad. Radula ( Figure 5 (f,i)) typical of Litthabitella . Penis ( Figure 9 (f)) with rather narrow tip and longer and broader filament than in the other Litthabitella species. Figure 2. Some of the studied localities: (a) Kostanje (locality 1); (b) Trsteno (locality 5); (c) Tomići (locality 7); (d) Sotonići, spring Voda u Križan (locality 8); (e,f) Levkas Island (locality 10). Distribution and habitat Known from the type locality only: Kostanje, a medium-sized spring at the right bank of the Cetina River ( Figure 2 (a)).