The subtribes and genera of the tribe Listroderini (Coleoptera, Curculionidae, Cyclominae): Phylogenetic analysis with systematic and biogeographical accounts Author Morrone, Juan J. Museo de Zoologia " Alfonso L. Herrera ", Departamento de Biologia Evolutiva, Facultad de Ciencias, Universidad Nacional Autonoma de Mexico (UNAM), Apartado Postal 70 - 399, 04510 Mexico D. F., Mexico juanmorrone2001@yahoo.com.mx text ZooKeys 2013 2013-02-28 273 15 71 http://dx.doi.org/10.3897/zookeys.273.4116 journal article http://dx.doi.org/10.3897/zookeys.273.4116 1313-2970-273-15 2D52FFD4495DFFF0FFE4FFDA0E63FE3F 578183 Listronotus Jekel, 1865 Fig. 21 Macrops Kirby, 1837: 199 ( non Wagler 1830, nec Burmeister 1835) (type species: not designated). Hyperodes Jekel, 1865: 566 (type species: Listroderes humilis Gyllenhal, 1834, by original designation). Listronotus Jekel, 1865: 566. Anchodemus LeConte, 1876: 181 (type species: Anchodemus hubbardi LeConte, 1876, subsequent designation by Kuschel 1950 : 14). Lixellus LeConte, 1876: 182 (type species: Lixellus filiformis LeConte, 1876, by indication, monotypy). Mascarauxia Desbrochers des Loges, 1898: 52 (type species: Mascarauxia cyrtica Desbrochers des Loges 1898, by indication, monotypy). Relistrodes Brethes , 1910: 209 (type species: Relistrodes breyeri Brethes , 1910, by indication, monotypy). Aulametopiellus Brethes , 1926: 415 (type species: Aulametopiellus dauci Brethes , 1926, by indication, monotypy). Mascaranxia Bosq, 1935: 330 (lapsus). Pseudhyperodes Hustache, 1939a: 49 (type species: Pseudhyperodes elongatus Hustache, 1939). Type species. Rhynchaenus caudatus Say, 1824 (subsequent designation by Henderson 1940 ). Diagnosis. Very small to medium-sized (1.0-14.0 mm); vestiture of subcircular scales and setae; antennal insertion distal; funicular segment 1 subequal to or shorter than 2; postocular lobes present, well-developed; elytra oblong-oval to elongate-oval, with intervals convex. Relationships. Listronotus is the sister genus to Steriphus (Australia). In a previous analysis based only on American taxa ( Morrone 1997a ), Listronotus was considered to be the sister genus to Neopachytychius . Species included. Listronotus alternatus (Dietz, 1889); Listronotus americanus LeConte, 1876; Listronotus angustatus (Champion, 1902); Listronotus annulipes (Blatchley, 1925); Listronotus anthracinus (Dietz, 1889); Listronotus apicalis (Hustache, 1926); Listronotus appendiculatus (Boheman, 1842); Listronotus argentinensis (Hustache, 1926); Listronotus arizonicus O'Brien , 1981; Listronotus blandus Henderson, 1940; Listronotus blatchleyi Henderson, 1940; Listronotus bonariensis (Kuschel, 1955); Listronotus borrichiae O'Brien , 1981 ; Listronotus bosqi (Hustache, 1926); Listronotus breyeri ( Brethes , 1910); Listronotus burkei O'Brien , 1981; Listronotus californicus (Dietz, 1889); Listronotus callosus LeConte, 1876; Listronotus carinatus (Blatchley, 1928); Listronotus carinicollis (Hustache, 1926); Listronotus caudatus (Say, 1824); Listronotus cinnamoneus (Hustache, 1926); Listronotus conabilis O'Brien , 1981; Listronotus crypticus O'Brien , 1981; Listronotus cryptops (Dietz, 1889); Listronotus cyrticus (Desbrochers des Loges, 1898); Listronotus dauci ( Brethes , 1926); Listronotus debilis Blatchley, 1916; Listronotus deceptus (Blatchley, 1916); Listronotus delumbis (Gyllenhal, 1834); Listronotus dietrichi (Stockton, 1963); Listronotus dietzi O'Brien , 1979; Listronotus distinctus Henderson, 1940; Listronotus dorsalis (Dietz, 1889); Listronotus dorytomoides (Hustache, 1926); Listronotus durangoensis O'Brien , 1977; Listronotus echinatus (Dietz, 1889); Listronotus echinodori O'Brien , 1977; Listronotus elegans Van Dyke, 1929; Listronotus elegantulus O'Brien , 1981; Listronotus elongatus (Hustache, 1939); Listronotus fasciatus O'Brien , 1981; Listronotus filiformis (LeConte, 1876); Listronotus frontalis LeConte, 1876; Listronotus geminatus (Hustache, 1926); Listronotus griseus (Hustache, 1926); Listronotus grypidioides (Dietz, 1889); Listronotus haldemani (Burke, 1963); Listronotus hirtellus (Dietz, 1889); Listronotus hoodi (Stockton, 1963); Listronotus hornii (Dietz, 1889); Listronotus hubbardi (LeConte, 1876); Listronotus humilis (Gyllenhal, 1834); Listronotus hyperodes (Dietz, 1889); Listronotus incompletus (Hatch, 1971); Listronotus ingens Henderson, 1940; Listronotus insignis Henderson, 1940; Listronotus laevis (Hustache, 1926); Listronotus laramiensis (Angell, 1893); Listronotus latinasus (Blatchley, 1922); Listronotus lineolaticollis (Blanchard, 1851); Listronotus lodingi (Blatchley, 1920); Listronotus lucens (Hustache, 1926); Listronotus lutulentus (Boheman, 1843); Listronotus maculatus (Hatch, 1971); Listronotus maculicollis (Kirby, 1837); Listronotus manifestus Henderson, 1940; Listronotus marginalis O'Brien , 1977; Listronotus marginicollis (Hustache, 1926); Listronotus marshalli O'Brien , 1981; Listronotus meridionalis O'Brien , 1977; Listronotus minutus (Blanchard, 1851); Listronotus montanus (Dietz, 1889); Listronotus nebulosus LeConte, 1876; Listronotus neocallosus O'Brien , 1981; Listronotus nevadicus LeConte, 1876; Listronotus nigropunctatus (Suffrian, 1871); Listronotus novellus (Blatchley, 1916); Listronotus obscurellus (Dietz, 1889); Listronotus obtectus (Dietz, 1889); Listronotus oregonensis (LeConte, 1876); Listronotus ornatipennis (Blanchard, 1851); Listronotus pallidus O'Brien , 1981; Listronotus palustris Blatchley, 1916; Listronotus pampaensis (Voss, 1954); Listronotus peninsularis (Blatchley, 1916); Listronotus plumosiventris O'Brien , 1977; Listronotus porcellus (Say, 1831); Listronotus poseyensis (Blatchley, 1916); Listronotus pseudosetosus O'Brien , 1981; Listronotus puncticollis (Hustache, 1926); Listronotus punctiger LeConte, 1876; Listronotus pusillus (Hustache, 1926); Listronotus rotundicollis LeConte, 1876; Listronotus rubtzoffi O'Brien , 1981; Listronotus rufomarginatus (Hustache, 1939); Listronotus salicorniae O'Brien , 1981; Listronotus scapularis Casey, 1895; Listronotus setosipennis (Hustache, 1926); Listronotus setosus LeConte, 1876; Listronotus similis Henderson, 1940; Listronotus sondondoanus (Voss, 1954); Listronotus sordidus (Gyllenhal, 1834); Listronotus sparsus (Say, 1831); Listronotus squamiger (Say, 1831); Listronotus sulcipennis (Boheman, 1834); Listronotus suturalis O'Brien , 1981; Listronotus teretirostris (LeConte, 1857); Listronotus testaceipes (Champion, 1902); Listronotus texanus (Stockton, 1963); Listronotus truncatus (Hatch, 1971); Listronotus tuberosus LeConte, 1876; Listronotus turbatus O'Brien , 1981; Listronotus vitticollis (Kirby, 1837); Listronotus vulgaris (Hustache, 1926); Listronotus wallacei (Stockton, 1963); Listronotus weiseri (Hustache, 1926). Host plants. Listronotus appendiculatus : Sagittaria latifolia Willdenow ( Alismataceae ); Listronotus argentinensis : Triticum aestivum L. ( Poaceae ); Listronotus blandus : Polygonum hydropiperoides Michx. ( Polygonaceae ); Listronotus bonariensis : Dactylis glomerata L., Festuca arundinacea Schreber, Hordeum vulgare L., Lolium multiflorum L., Lolium perenne L., Poa annua L., Triticum aestivum L., Zea mays L. ( Poaceae ) and Trifolium repens L. ( Fabaceae ); Listronotus borrichiae : Borrichia frutescens (L.) DC ( Asteraceae ) and Salvinia sp. ( Salviniaceae ); Listronotus caudatus : Polygonum bicorne Raf. ( Polygonaceae ); Listronotus cinnamoneus : Limnobium stoloniferum (G. F . W. Meyer) Griseb. ( Hydrocharitaceae ); Listronotus cryptops : Sagittaria lancifolia L. ( Alismataceae ); Listronotus dauci : Daucus carota L. ( Apiaceae ); Listronotus dietrichi : Dahlia sp. ( Asteraceae ), Gossypium sp. ( Malvaceae ), Persus sp. ( Lauraceae ), Phaseolus sp. ( Fabaceae ), Cenchrus sp., Chloris sp., Cynodon sp., Eleusine sp., Zea sp. ( Poaceae ), Coffea sp. ( Rubiaceae ), Lycopersicum sp. ( Solanaceae ) and Menta sp. ( Lamiaceae ); Listronotus echinodori : Echinodorus cordifolius (L.) Griseb. and Sagittaria latifolia Willdenow ( Alismataceae ); Listronotus elongatus : Hydrocotyle ranunculoides L. f. ( Apiaceae ); Listronotus haldemani : Juncus nodatus Coville in N. L. Britton and A. Brown ( Juncaceae ); Listronotus maculicollis : Agrostis palustris Huds. and Poa annua L. ( Poaceae ); Listronotus manifestus : Sagittaria longiloba Engelm. ex J. G. Sm. ( Alismataceae ); Listronotus marginicollis : Myriophyllum aquaticum (Velloso) Verde ( Haloragaceae ); Listronotus montanus : Triticum aestivum L. ( Poaceae ); Listronotus neocallosus : Sagittaria engelmanniana J. G. Smith, Sagittaria graminea Michaux and Sagittaria stagnorum Small ( Alismataceae ); Daucus carota L. and Petroselinum crispum (Miller) A. W. Hill. ( Apiaceae ) ( Listronotus oregonensis ); Listronotus plumosiventris : Sagittaria latifolia Willdenow ( Alismataceae ); Listronotus rotundicollis : Crinum sp. ( Amaryllidaceae ); Listronotus rubtzoffi : Sagittaria cuneata Sheldon ( Alismataceae ); Listronotus salicorniae : Salicornia virginica L. ( Amaranthaceae ); Listronotus scapularis : Sagittaria longiloba Engelm. ex J. G. Sm. and Sagittaria sp. ( Alismataceae ); Listronotus setosipennis : Parthenium hysterophorus L. ( Asteraceae ); Listronotus similis : Paspalum distichum L. ( Poaceae ) and Polygonum bicorne Raf. ( Polygonaceae ); Listronotus teretirostris : Eleocharis macrostachya Britton ( Cyperaceae ); Listronotus texanus : Daucus carota L. ( Apiaceae ); Listronotus turbatus : Sagittaria sp. ( Alismataceae ) ( Brethes 1926 ; Burke 1963 ; Martel et al. 1976 ; May 1977 ; O'Brien 1977 , 1981; Cordo and DeLoach 1982 ; Cordo et al. 1982 ; Edelson 1985 ; Barker 1989; Boivin et al. 1990 ; Kuschel 1990 ; Maes and O'Brien 1990 ; Anderson 1992 ; Wild et al. 1992 ; May 1993 ; Cragnolini 1994 ; Blodgett et al. 1997 ; Lanteri et al. 2002 ; Torres and Casey 2002 ; Rothwell 2003 ). Immature stages. Listronotus bonariensis (May, 1977, 1993, 1994; Marvaldi, 1998). Geographical distribution. Widespread in the Americas, from Canada to Argentina and Chile ( O'Brien 1977 , 1981; O'Brien and Wibmer 1982 ; Wibmer and O'Brien 1986 ; Maes and O'Brien 1990 ; Anderson 1992 ). This distribution corresponds to the Nearctic, Neotropical and Andean regions, as well as the South American and Mexican Transition Zones. Material examined. Listronotus americanus (BMNH), Listronotus apicalis (MLP), Listronotus appendiculatus (AMNH, BMNH), Listronotus argentinensis (AMNH, MACN, MLP9, Listronotus blandus (AMNH, BMNH), Listronotus bonariensis (BMNH, MHNS), Listronotus bosqi (BMNH, MLP, MZFC), Listronotus breyeri (MACN, MZFC), Listronotus californicus (AMNH), Listronotus callosus (BMNH, AMNH), Listronotus caudatus (BMNH, AMNH), Listronotus cinnamoneus (MLP), Listronotus cryptops (BMNH, AMNH), Listronotus cyrticus (AMNH, MACN, MLP), Listronotus dauci (MACN), Listronotus debilis (AMNH), Listronotus delumbis (BMNH, AMNH), Listronotus dietzi (AMNH), Listronotus distinctus (BMNH), Listronotus durangoensis (AMNH, BMNH), Listronotus echinatus (AMNH), Listronotus echinodori (AMNH, BMNH), Listronotus elongatus (MLP, MZFC), Listronotus filiformis (BMNH, AMNH), Listronotus frontalis (AMNH, BMNH), Listronotus geminatus (MACN, MLP), Listronotus griseus (AMNH, MACN, MLP), Listronotus grypidioides (AMNH), Listronotus haldemani (BMNH), Listronotus hornii (AMNH), Listronotus hubbardi (BMNH), Listronotus humilis (AMNH), Listronotus hyperodes (AMNH), Listronotus incompletus (AMNH), Listronotus ingens (AMNH), Listronotus lineolaticollis (MLP), Listronotus lutulentus (BMNH), Listronotus maculicollis (AMNH), Listronotus manifestus ( AMNH, BMNH), Listronotus marginalis (BMNH), Listronotus marginicollis (MACN, MLP), Listronotus meridionalis (BMNH), Listronotus minutus (AMNH), Listronotus nebulosus (AMNH), Listronotus novellus (AMNH), Listronotus oregonensis (AMNH, BMNH, MZFC), Listronotus ornatipennis (MHNS), Listronotus palustris (AMNH, BMNH), Listronotus plumosiventris (BMNH), Listronotus porcellus (AMNH), Listronotus puncticollis (MLP), Listronotus punctiger (AMNH, BMNH), Listronotus pusillus (MLP, MZFC), Listronotus rotundicollis (AMNH, BMNH), Listronotus rubtzoffi (AMNH), Listronotus rufomarginatus (MLP), Listronotus scapularis (AMNH), Listronotus setosipennis (MLP), Listronotus setosus (AMNH), Listronotus similis (AMNH, BMNH), Listronotus sordidus (AMNH, BMNH), Listronotus sparsus (AMNH, BMNH), Listronotus squamiger (AMNH, BMNH), Listronotus teretirostris (AMNH, BMNH), Listronotus texanus (AMNH), Listronotus truncatus (AMNH), Listronotus tuberosus (AMNH, BMNH), Listronotus vitticollis (AMNH) and Listronotus vulgaris (MLP).