A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands Author Xu, Xin Author Ono, Hirotsugu Author Kuntner, Matjaz Author Liu, Fengxiang Author Li, Daiqin text ZooKeys 2019 888 1 50 http://dx.doi.org/10.3897/zookeys.888.34494 journal article http://dx.doi.org/10.3897/zookeys.888.34494 1313-2970-888-1 B995C05697EC41A49012B58F9D3AFDC1 F8810409F4DA5A43BF94417F5D40DECE Heptathela tokashiki sp. nov. Fig. 21 Type material. Holotype : JAPAN · ♀; Okinawa-ken, Tokashikijima Island, Aharen; 26.19N , 127.37E ; alt. 100 m; 8 May 2014; D. Li and B. Wu leg.; XUX-2014-062. Paratypes : JAPAN · 25 ♀♀; same data as for holotype; XUX-2014-046 to 063D · 3 ♀♀; same data as for holotype; 27 December 2012; D. Li, F.X. Liu and X. Xu leg.; XUX-2012-417, 421, 425. Diagnosis. Females of H. tokashiki sp. nov. can be distinguished from H. shuri sp. nov. by the long inner receptacular clusters with several granules ( Fig. 21 A-D ). Heptathela tokashiki sp. nov. can also be diagnosed from all other Okinawa group Heptathela species by the following unique nucleotide substitutions in the standard DNA barcode alignment: T (68), A (200), C (290), A (362). Figure 21. Female genital anatomy of Heptathela tokashiki sp. nov. A, B 4062 (holotype, short for XUX-2014-062) C, D 4063 A, C vulva dorsal view B, D vulva ventral view. Scale bar: 0.5 mm. Description. Female (Holotype). Carapace brown; opisthosoma dark brown, with dark brown tergites close to each other; cheliceral groove with eleven pronounced denticles; seven spinnerets. Measurements: BL 11.80, CL 5.15, CW 4.10, OL 6.80, OW 5.10; ALE> PLE> PME> AME; palp 8.60 (3.05 + 1.50 + 1.95 + 2.10), leg I 10.50 (3.35 + 1.85 + 2.15 + 2.10 + 1.05), leg II 10.45 (3.40 + 1.75 + 2.00 + 2.20 + 1.10), leg III 10.95 (3.20 + 1.85 + 1.80 + 2.60 + 1.50), leg IV 15.70 (4.60 + 2.00 + 2.80 + 3.90 + 2.40). Female genitalia . A pair of depressions on the ventro-lateral part of the genital atrium ( Fig. 21B, D ). Paired receptacular clusters along the anterior margin of bursa copulatrix, divided into two parts, the inners longer than the laterals, both with several granules, without genital stalks ( Fig. 21A, B ). Male. Unknown. Etymology. The species epithet, a noun in apposition, refers to the type locality. Distribution. The species is endemic to the Japanese island Tokashikijima ( Fig. 1C ).