World reclassification of the Cardiophorinae (Coleoptera, Elateridae), based on phylogenetic analyses of morphological characters Author Douglas, Hume B. text ZooKeys 2017 655 1 130 http://dx.doi.org/10.3897/zookeys.655.11894 journal article http://dx.doi.org/10.3897/zookeys.655.11894 1313-2970-655-1 8C475FAB25E044CEA2FBC3B83F316D8C 8C475FAB25E044CEA2FBC3B83F316D8C Cardiodontulus Van Zwaluwenburg, 1963 Figs 147-150 Cardiodontulus Van Zwaluwenburg, 1963: 341. Type species: Cardiodontulus brandti Van Zwaluwenburg, 1963: 341. Diagnosis. Prothorax. Pronotum with lateral carina extending anterad from hind angles following lateral edge, not reaching anterior edge; procoxal cavities closed. Pterothorax. Scutellum with middle of anterior edge concave: broadly or abruptly emarginate. Legs. Tarsomere 4 lobed or lamellate; Tarsal claws with 2 points per side (Fig. 150). Also. Apical half of elytral interval 9 not costate. Known from: Papua New Guinea, Australia. Only males known, 2 spp.