Taxonomic notes on some Cantharidae of Sicily and North Africa
Author
Fanti, Fabrizio
text
Baltic Journal of Coleopterology
2020
2020-12-30
20
2
141
152
journal article
294381
10.5281/zenodo.10905414
d953bb02-00f3-494c-a385-e00ea60c127b
1407-8619
10905414
Ancistronycha neapolitana
(
Pic, 1918
)
comb. nov. et stat. nov.
Cantharis Erichsoni
v. nov.
neapolitanus
Pic, 1918: 22
.
Loc. typ
.: “Environs de Naples (Pic)”
[
Ancistronycha
is a feminine name therefore the correct declination proposed here is
neapolitana
,
declinatio nov.]
=
Cantharis sicula
Pic, 1906: 197-198
.
Loc. typ.
: “
Sicile
(communiqué par
Mr. E. Ragusa
et aussi dans ma collection)”
[
syn. nov.
] [junior homonym of
Cantharis livida
var.
sicula
Bourgeois, 1893: 17
]
=
Cantharis lucens
Moscardini, 1967: 30
(aedeagus drawings - caption), 32-33 [
syn. nov.
]
=
Cantharis lucens
f.
fumosothorax
Moscardini, 1967: 33
“[
syn. nov.
]”. [Name not available because it dates back to 1961 (
ICZN
1999Art. 45.6.3.)]
PORTA 1929: 56
, 56 note [
C. sicula
];
MOSCARDINI 1967: 29
, 30
Fig. 1
–
cd
[aedeagus, as
C. lucens
], 31 [distribution map], 1968: 55 [localities, as
C. lucens
];
KAZANTSEV 2005
(as
A. lucens
): 205 [aedeagus], 207 [key];
FANTI 2014: 64
(as
A. lucens
), 70 (as
C. sicula
).
Cantharis sicula
Pic (1906)
was found to be a junior homonym of
Cantharis livida
var.
sicula
Bourgeois, 1893
(see also
Fiori 1914: 60
) and therefore is a name that’s not available (
ICZN 1999
Art. 57.2.). While the name of Bourgeois was described before 1961 and, without any ambiguity, as a variety (“var.”), it must be considered of subspecific rank and therefore available (
ICZN 1999
Art. 45.6.4.). Furthermore, in addition to this validity, it should be noted that the Bourgeois species was sometimes considered a “subspecies” (
e.g.
,
Portevin 1931: 407
[indicated as a variety but intuitively should be considered a subspecies]) before 1985 (
ICZN 1999
Art. 45.6.4.1.), and sometimes even more recently (
Liberti 1995
). Therefore, it is unclear what name should be used for the species
C. sicula
Pic.
The study of the
C. sicula
holotype
allowed me to attribute it, as had already been suggested by
Fanti (2014)
, to the genus
Ancistronycha
Märkel.
The specimen is a female, and has an evident talon-shaped tooth on the claw in the front legs and a pronotum with slightly rounded sides. Comparing it with the specimens of
Ancistronycha lucens
f.
fumosothorax
from the Fiori collection in the Experimental Evolutionary Biology Department of the University of Bologna and also based on work by
Moscardini (1967)
, it is possible to determine that it is the same species. This aspect was, in a very small part, also suggested by Fiori himself (1914: 57, 59-60); however, he still described them as
Cantharis puncticollis
Levrat.
Fig. 5.
Ancistronycha neapolitana
(
Pic, 1918
)
. Specimens of Calabria (
“var.
humeralis
” mihi).
Fig. 6.
Metacantharis puncticollis
(
Levrat, 1857
)
. Neotypus preserved in the Walter Wittmer collection at the Natural History Museum of Basel.
Fig. 7.
Metacantharis puncticollis
(
Levrat, 1857
)
. Original label of Neotypus.
When studying the literature of the
Ancistronycha
varieties ascribable to
C. sicula
and/or to the related
A. erichsonii
(Bach, 1852)
present in Central-Southern
Italy
and
Sicily
and which may be the first available name, we first find a
variety described
as
inapicalis
(
Pic 1902: 63
sub
Cantharis Erichsoni
variété
inapicalis
), which theoretically could take precedence. This variety is, however, described without indication of locality (
holotype
unobtainable and probably no longer preserved), which remains unknown. In the absence of the type and other information, we can only vaguely hypothesize that it comes from
Piedmont
/ North-West
Italy
(
ICZN 1999
Recommendation 76A.), since the collector is Baudi and most of his collections are from those areas. Therefore, it must still be considered as a form of
A. erichsonii
(
Kazantsev & Brancucci 2007
)
. Later, we find described from the surroundings of Naples, the
variety
neapolitanus
(
Pic 1918: 22
sub
Cantharis Erichsoni
v. nov.
neapolitanus
), whose
holotype
has not been located at present (Taghavian-Azari personal communication) but which could still be present and mixed with other species of the
Maurice
Pic collection. This
variety
neapolitanus
is, however, clearly related to
C. sicula
(no other species with this coloration are present in the type locality. Furthermore,
Cantharis livida
and
Rhagonycha fulva
are different) and therefore is the first name available and should be used instead. Finally, Moscardini described the species
Cantharis lucens
, which becomes a synonym (
Moscardini 1967
).
Fig. 8.
Metacantharis puncticollis
(
Levrat, 1857
)
. Overview of the specimens from the Wittmer collection in Basel.
Note.
Ancistronycha neapolitana
is an Italian endemic species (though there is a citation of
A. erichsonii
from Dalmatia that
Moscardini, 1968
refers to as this species), that is rather rare and is found in medium-high altitudes on Umbelliferae from Emilia-Romagna to Sicily (
Moscardini 1967: 32-33
;
Fanti 2014: 64
and 70), and with only a specimen known from 200 meters above sea level (present work). The northern limit is not well known but it is currently considered the area of the Orrido di Botri, Minucciano, San Godenzo (present work) and Camaldoli in Tuscany, and Fanano in Emilia-Romagna (present work, and first citation for Emilia-Romagna). In the Ligurian Apennines: Torriglia, Monte Penna (
Moscardini 1967
), and in the Alps and Prealps, it is replaced by the related
A. erichsonii
(Bach)
, which is a species from Central Europe,
Romania
,
Ukraine
,
Georgia
and
Turkey
. In the Apennine,
A. neapolitana
has a head, pronotum and legs testaceous with black tarsi (sometimes first tarsomere of the front legs is two-colored: black and testaceous), and has an elytra with black apex that extends to the elytral half (
two females
from Calabria mentioned in this work, have black elytra with only the testaceous humeri, of which in one of these the testaceous part covers a very little portion of the humeri themselves.
Fig. 5
). In Sicily, where only
four specimens
are currently known (coll. Pic:
1 female
and coll. Fiori:
1 male
2 females
), it is characterized by a head behind the eyes that is brown-piceous and by a brown-piceous pronotum with more or less broadly yellowish sides, with yellowish testaceous legs and fully testaceous elytra or with the blackish apex up to about half of its length.