Systematic revision of the trilobite genera Laudonia and Lochmanolenellus (Olenelloidea) from the lower Dyeran (Cambrian Series 2) of western Laurentia Author Webster, Mark Author Bohach, Lisa L. text Zootaxa 2014 3824 1 1 66 journal article 36828 10.11646/zootaxa.3824.1.1 df227efc-13d9-421e-92ed-f9aa417b0cf7 1175-5326 286404 023D78D0-4182-48D2-BAEB-CDA6473CF585 Family “ Laudoniidae ” Palmer & Repina, 1993 The subfamily Laudoniinae Palmer & Repina, 1993 was originally established to include the genera Laudonia and (tentatively) Olenelloides Peach, 1894 ( Palmer & Repina, 1993 ; although if Olenelloides was placed in the same subfamily as Laudonia , then Olenelloidinae Hupé, 1953 would have priority over Laudoniinae). Lieberman (1998) subsequently reassigned some material of Laudonia to a new genus Lochmanolenellus . The cladistic analysis presented by Lieberman (1998) found that both Lochmanolenellus and Olenelloides shared more recent common ancestry with Bristolia , Fremontella Harrington, 1956 , Nephrolenellus , Bolbolenellus Palmer and Repina, 1993 , Peachella , and Biceratops than with Laudonia bispinata , so Lochmanolenellus and Olenelloides were accordingly assigned to the Biceratopsidae Pack & Gayle, 1971 . Laudonia bispinata was determined to be the immediate sistertaxon to the Biceratopsidae , with Mummaspis occidens and Fritzolenellus truemani forming successive outgroups to the [ Laudonia bispinata + Biceratopsidae ] clade ( Lieberman, 1998 ). For convenience, Lieberman (1998) grouped Laudonia , Mummaspis Fritz, 1992 , and Fritzolenellus Lieberman, 1998 into a paraphyletic “ Laudoniidae ”. Lieberman’s (1998) cladistic analysis is fraught with problems (e.g., character coding issues discussed following; similar coding problems have been identified in other taxa [ Webster, 2007a , 2009 ]). However, a revised analysis of the phylogenetic affinities of Laudonia is beyond the scope of the present paper, and Lieberman’s (1998) convention is followed herein.