The oldest fossil of the family Issidae (Hemiptera, Fulgoromorpha) from the Paleocene of Menat (France) Author Bourgoin, Thierry Author Wang, Menglin Author Nel, André text European Journal of Taxonomy 2020 2020-01-22 596 1 8 journal article 24115 10.5852/ejt.2020.596 bb0a8a5e-6a00-4b09-90e7-e7fc2c73bb2c 3659729 28C76456-50AA-4DD7-A143-052433A51DD2 Cubicostissus palaeocaeni Bourgoin & Wang sp. nov. urn:lsid:zoobank.org:act: 3E61A3B6-E035-41B1-96CD-B450B0F71F93 Figs 1–2 Diagnosis Described from a single imprint from Menat ( Fig. 1 ). Separated from all Hysteropterini species by its elongated forewing, the narrow postcostal band and the conformation of the venation. Etymology The specific epithet means ‘from the Palaeocaenum (Paleocene)’ and refers to the age of the deposit where the fossil was discovered. Material examined Holotype FRANCE • 1 specimen ; Puy-de-Dôme , volcano-sedimentary maar of Menat ; Paleocene , Selandian ; Musée de la Paléontologie , Menat ; NEL 3485. Description Forewing ( Figs 1–2 ) 2.6 times as long as wide, wider before mid-length: 4.04 mm long, 1.63 mm wide before mid-length at MP forking level. Basal cell (bc) short, 0.29 mm long. No hypocostal plate. Anterior margin slightly and regularly convex, maximum convexity at level of MP fork. Posterior margin slightly concave at the end of the clavus. Common stem ScP+R more than twice as long as basal cell ( 0.67 mm long). Postcostal cell forming a narrow band, slightly narrower than radial cell, itself narrower than basal cell. ScP+R forking late: ScP+R 2.7 times as long as basal cell. Radial cell open, as wide as C1 cell before mid-length of tegmen, with ScP+RA and RP running parallel. Vein MP 1.33 mm long, forking late in cell C3 at same level of fusion of Pcu and A1; MP1+ 2 in straight line with vein MP; MP3+4 diverging posteriorly. CuA almost straight, single up to apical level of clavus, separated from MP and MP3+4 by a wide area (open median cell) almost as wide as area width between MP+MP3+4 and costal margin. C5 cell open. CuA connected with parallel CuP before mid-length of tegmen by four transverse veins. Just before reaching clavus, apically forked with CuA1 starts running parallel to the tegmen margin and CuA2 joining tegmen margin. Pcu and A1 connecting late in the last quarter of a long closed clavus. A1 closely parallel to posterior margin. Traces of brownish coloration and small micro-granulations are visible in cubito-anal and subcostal areas ( Fig. 1 ).