The oldest fossil of the family Issidae (Hemiptera, Fulgoromorpha) from the Paleocene of Menat (France) Author Bourgoin, Thierry Author Wang, Menglin Author Nel, André text European Journal of Taxonomy 2020 2020-01-22 596 1 8 journal article 24115 10.5852/ejt.2020.596 bb0a8a5e-6a00-4b09-90e7-e7fc2c73bb2c 3659729 28C76456-50AA-4DD7-A143-052433A51DD2 Genus Cubicostissus Bourgoin & Nel gen. nov. urn:lsid:zoobank.org:act: 3F1F9C1A-4698-4BFA-9541-2CFC3E5EECCB Type species Cubicostissus palaeocaeni sp. nov. , designated here by monotypy. Diagnosis Cubicostissus gen. nov. separates from other Hysteropterini genera by their forewing more than 2.5 as long as wide, wider before mid-length, with anterior margin slightly and regularly convex and posterior margin slightly concave at the end of the clavus. The postcostal cell forms a narrow band, slightly narrower than the radial cell band, and as wide as C1 cell. ScP+RA and RP fork late with ScP+R more than 2.5 times as long as basal cell. MP forks in C3 at the same level of Pcu and A1 fusion and CuA remains single up to the apical level of the clavus, then forks in an open C5. CuP is almost straight, connected by transverse veins to CuA. Pcu and A1 are connected in the last quarter of the clavus. Etymology Arbitrary combination referring to the forewing CuA vein ( cubito costa ) single and the generic name Issus . Note Previously placed in Issinae sec. Wang et al. (2016) , Hysteropterini were recently regarded deserving a subfamily rank ( Zhao et al. 2019 ). They differ from Issinae Issini by their late forking of CuA after the end of the clavus while CuA forks well before in Issini genera. By its venation schema, Cubicostissus gen. nov. tegmen approaches species of Tshurtshurnella Kusnezov, 1927 , but it appears more elongated: 2.6 times as long as wide (× 2 or less in Tshurtshurnella ) and with a late fusion of Pcu and A 1 in the last quarter of the clavus (earlier in Tshurtshurnella ). From Hysteropterum Amyot & Audinet-Serville, 1843 , it differs by the elongated tegmen, the absence of identified apical cells, the presence of some transverse veins between CuA and CuP and, at mid-length, the costal area almost as wide as C1 and the open radial cell. Its simpler venation also easily separates this new taxon from all the other issid fossils currently known.