Taxonomic review of the genus Lista Walker, 1859 from China (Lepidoptera, Pyralidae, Epipaschiinae), with descriptions of five new species Author Rong, Hua College of Life Sciences, Nankai University, Tianjin, China Author Wang, Yiping College of Life Sciences, Nankai University, Tianjin, China Author Qi, Mujie 0000-0002-8953-3422 College of Life Sciences, Nankai University, Tianjin, China & lihouhun @ nankai. edu. cn; https: // orcid. org / 0000 - 0002 - 8953 - 3422 lihouhun@nankai.edu.cn Author Li, Houhun 0000-0002-1931-8401 College of Life Sciences, Nankai University, Tianjin, China & qimujie @ 163. com; https: // orcid. org / 0000 - 0002 - 1931 - 8401 text Zootaxa 2021 2021-12-10 5081 2 237 262 journal article 3019 10.11646/zootaxa.5081.2.4 24015f76-6d6f-461f-be58-aa9ff6ae55ed 1175-5326 5771739 4AABAEE0-115D-4316-867E-F16E1125C2C2 Lista insulsalis ( Lederer, 1863 ) ( Figs 12 , 27 , 42 ) Paracme insulsalis Lederer, 1863: 339 . TL: China ( Zhejiang ). Stericta rubiginetincta Caradja, 1925: 314 . TL: China ( Guangdong ). Lista insulsalis (Lederer) : Solis, 1992: 283 . Lista haraldusalis ( Walker, 1859 ) : Li, Ren et al. , 2009: 35 , misidentified. Material examined. CHINA , Tianjin : 2 ♀ , Mt. Baxian , 24.VII.2015 , coll. HH Li and PX Cong ; Hebei Province : 1 ♀ , Jingxing County , 27.VII.1998 ; Zhejiang Province : 1 ♀ , Chanyuansi , Mt. Tianmu , 350 m , 15.VIII.1999 , coll. HH Li et al .; 1 ♀ , Xianrending , Mt. Tianmu , 1500 m , 18.VIII.1999 , coll. HH Li et al .; 8 ♂ , 4 ♀ , Mt. Fengyang , Longquan , Lishui , 1470 m , 26–31.VII.2007 , coll. Q Jin ; 1 ♀ , Wuyanling , Taishun , 790 m , 2.VIII.2007 , coll. Q Jin ; 1 ♀ , Mt. Jiulong , 4.VIII.2011 , coll. XB Fu ; 1 ♂ , 1 ♀ , Xianrending , Mt. Tianmu , 1500 m , 25.VII.2011 , coll. LL Yang and N Chen ; 1 ♂ , Zhangkengkou , Mt. Tianmu , 623 m , 5.VII.2013 , coll. AH Yin and XC Wang ; 3 ♂ , 3 ♀ , Huangtanyu, Mt. Jiulong , 467 m , 6–9.VII.2013 , coll. AH Yin and XC Wang ; 3 ♂ , 1 ♀ , Neijiujian , Mt. Jiulong , 430 m , 7.VII.2013 , coll. AH Yin and XC Wang ; 4 ♂ , 3 ♀ , Tianmu Village , Mt. Tianmu , 335 m , 1.VII.2014 , coll. AH Yin et al .; 1 ♂ , Qianqingtang , Qingliangfeng , 1059 m , 28.VII.2014 , coll. AH Yin et al .; 1 ♂ , Sanmuping , Mt. Tianmu , 789 m , 17.VII.2015 , coll. AH Yin et al .; 4 ♂ , 3 ♀ , Laofoyan Village , Shuangxikou , Jiangkou , 424 m , 7–8.VIII.2016 , coll. QY Wang et al ., slide Nos. RH16173 ♂ , RH16174 ♀ ; 1 ♀ , Huangtianhu Forest Farm, Jingning , 787 m , 11.VIII.2016 , coll. QY Wang et al .; 6 ♂ , 1 ♀ , Baishanzu Nature Reserves , Qingyuan , 1149 m , 13.VIII.2016 , coll. QY Wang et al .; 1 ♀ , Qiansiyan , Qingtian , 75 m , 18.VIII.2016 , coll. QY Wang et al .; 34 ♂ , 5 ♀ , Shimendong , Qingtian , 102 m , 20–21.VIII.2016 , coll. QY Wang et al ., slide Nos. RH16179 ♂ , RH16180 ♀ , RH16181 ♂ ; 8 ♂ , 5 ♀ , Linkeng Village , Yongjia , 387 m , 23–26.VIII.2016 , coll. QY Wang et al .; Anhui Province : 1 ♀ , Huangshan , 3.VII.2004 , coll. JS Xu and JL Zhang ; 1 ♀ , Ke Village , Mt. Jiuhua , 8.VIII.2004 , coll. JS Xu and JL Zhang ; Fujian Province : 1 ♀ , Mt. Tianzhu , Xiamen , 43 m , 15.VII.2010 , BB Hu and J Zhang ; 14 ♀ , Mt. Tianzhu , Xiamen , 220 m , 7–19.IX.2010 , coll. YH Sun and J Zhang ; 1 ♂ , Daiyun Village , Mt. Daiyun , 902 m , 23. V .2012, coll. JW Li ( SYSBM ); 1 ♀ , Tongmu Village , Wuyishan , 2.IX.2012 , coll. ZB Wang ; 4 ♀ , Mt. Tianzhu , 13–21.VII.2014 , coll. XF Yang ; Jiangxi Province : 1 ♂ , Mt. Sanqing , Dengxing County , 1550 m , 13.VIII.1985 ; 1 ♀ , Ganzhou Forestry Institute , 16.VII.2006 , coll. JS Xu and WC Li ; 1 ♂ , Wuyishan National Nature Reserves , 900 m , 13.VIII.2007 , coll. JS Xu ; Henan Province : 1 ♀ , Laojieling , 850 m , 17.VII.1998 , coll. ML Sheng ; 1 ♀ , Lingshansi , Mt. Luo , 350 m , 21. V .2000, coll. HL Yu ; 5 ♂ , 2 ♀ , Mt. Jigong , Xinyang , 700 m , 13.VII.2001 , coll. DD Zhang , slide No. WYP05012 ♂ ; 1 ♂ , 1 ♀ , Shuiliandong , Tongbai , 300 m , 16–17.VII.2001 , coll. DD Zhang ; 1 ♀ , Shiziping , Lushi , 1200 m , 19.VII.2001 , coll. DD Zhang ; 4 ♂ , Mt. Baiyun , Luoyang , 1560 m , 22.VII.2001 , coll. DD Zhang , slide No. WYP05041 ; 2 ♂ , Mt. Baiyun , Song County , 1580 m , 22.VII.2002 , coll. XP Wang ; 1 ♀ , Mt. Baiyun , Song County , 1400 m , 14.VIII.2008 , coll. HH Li et al .; Hubei Province : 1 ♂ , Shayuan , Hefeng , 1260 m , 15.VII.1999 , coll. HH Li et al .; 1 ♂ , 1 ♀ , Pingbaying , Xianfeng , 1280 m , 21.VII.1999 , coll. HH Li et al ., slide No. WYP05163 ♀ ; 1 ♀ , Bajiaomiao , Shennongjia , 1100 m , 18.VII.2003 , coll. SL Hao ; 3 ♂ , 3 ♀ , Mt. Wudang , Shiyan , 172 m , 6.IX.2012 , coll. JWi Li ( SYSBM ) ; 1 ♂ , Tiantangzhai , Luotian County , 570 m , 17.IX.2012 , coll. JW Li ( SYSBM ) ; 1 ♂ , Taohuachong , Mt. Dbie , 661 m , 24. VI .2014, coll. XH Chen and C Pan ( SYSBM ); 1 ♀ , Taohuachong , Yingshan County , 635 m , 26. VI .2014, coll. W Guan and MQ Yang ; 2 ♀ , Wujiashan , Yingshan County , 880 m , 30. VI .2014, coll. W Guan and MQ Yang ; 2 ♂ , 1 ♀ , Qingtaiguan , Mt. Dabie , 590 m , 1–3.VII.2014 , coll. LJ Xu ; 1 ♂ , Qingtaiguan Forest Farm, Luotian County , 570 m , 2.VII.2014 , coll. W Guan and MY Yang ; 1 ♀ , Qingtaiguan , Luotian County , 524 m , 2.VII.2014 , coll. ZH Liu and C Pan ( SYSBM ) ; Hunan Province : 1 ♀ , Xiaolang Village , Ketou , Xinhua County , 3.VIII.2004 , coll. YL Xiao ; 3 ♀ , Jianxin Village , Ketou , Xinhua County , 10–11.VIII.2004 , coll. YL Xiao , slide No. WYP05069 ; 1 ♂ , Zhupo Village, Huitong County , 23.VIII.2012 , coll. JW Li and XH Chen ( SYSBM ) ; Guangdong Province : 1 ♂ , Mt. Wutong , Shenzhen , 15.IX.1999 , coll. FL Jia ( SYSBM ) ; 1 ♂ , 1 ♀ , Heshan, 28. VI .2002 , coll. GL Liu ( SYSBM ); 1 ♀ , Dawuling Nature Reserves , Xinyi , 1000 m , 8.VIII.2003 , coll. DD Zhang ; 1 ♂ , Dawuling , Xinyi , 900 m , 11.VIII.2003 , coll. H Pang and BL Zhang ( SYSBM ) ; 1 ♀ , Yangmei , Gaoming , 23. VI .2006, coll. DD Zhang ( SYSBM ); 1 ♂ , 1 ♀ , Mt. Dadong , Lian County , 8.VII.2008 , coll. FX He ( SYSBM ) ; 2 ♂ , 1 ♀ , Heishiding , Fengkai County , 15. VI .2009, coll. FX He ( SYSBM ); 1 ♂ , 1 ♀ , Heishiding , Fengkai , 13.VIII.2010 , coll. HD Chen et al . ( SYSBM ); 1 ♀ , Heishiding, Fengkai , 2.VII.2011 , coll. B Tong and Y Li ( SYSBM ) ; 1 ♂ , Heishiding , Fengkai , 5.X.2011 , coll. B Tong and Y Li ( SYSBM ) ; 1 ♂ , Yanshuitian , Fengkai , 3. VI .2011, coll. HD Chen and B Tong ( SYSBM ); 1 ♀ , Yanshuitian , Fengkai , 3.VII.2011 , coll. B Tong and Y Li ( SYSBM ) ; 1 ♂ , Shimentai , Yingde , 27. V .2012, coll. LJ Yang and QJ Jia ( SYSBM ); 1 ♀ , Mt. Danxia , Shaoguan , 96 m , 6. VI .2012, coll. JW Li ( SYSBM ); 1 ♂ , 1 ♀ , Mt. Dinghu , Zhaoqing , 37 m , 15.X.2012 , coll. JW Li ( SYSBM ) ; 1 ♀ , Mt. Xiangtou , Huizhou , 67 m , 18.X.2012 , coll. JW Li ( SYSBM ) ; Guangxi Zhuangzu Autonomous Region : 2 ♂ , Hongqi Forest Farm, Shangasi , 260 m , 2.XI.2002 , coll. SL Hao and HJ Xue , slide No. WYP05011 ; 1 ♂ , Yongfu , Guilin , VI .2006 , coll. ZF Zhao ; 1 ♀ , Huaping , Yachang Forest Farm, Leye County , 910 m , 28.VII.2004 , coll. JS Xu ; 2 ♂ , 4 ♀ , Huaping , 950 m , 8.VIII.2006 , coll. WC Li ; 1 ♂ , Mt. Yuanbao , 700 m , 11.VIII.2006 , coll. WC Li ; 1 ♀ , Qingshan Forest Farm, Pingxiang , 300 m , 20.VII.2011 , coll. BB Hu et al . ; 1 ♂ , Mt. Daming , Nanning , 1200 m , 6.VIII.2011 , coll. SL Hao and YH Sun ; 1 ♂ , Shiwandashan , Shangsi , 22.VIII.2011 , coll. LJ Yang ( SYSBM ) ; 1 ♂ , Shiwandashan , Shangsi , 22.VIII.2011 , coll. DD Zhang ( SYSBM ) ; 1 ♂ , Shaoping Forest Farm, Pingxiang , 280 m , 20.IX.2012 , coll. ZG Zhang ; 1 ♀ , Dalongping , Cenwanglaoshan , 1290 m , 6.VIII.2014 , coll. XL Wei and C Ran ; 1 ♂ , 1 ♀ , Yinshan Park , Dayaoshan Nature Reserves , Jinxiu , 1364 m , 19–21.VII.2015 , coll. MJ Qi and SN Zhao ; Sichuan Province : 6 ♀ , Haichao , Luzhou , 25–30.VII.1995 , coll. YX Zeng , slide Nos. WYP05066, WYP05067 ; 1 ♀ , Nuoshuihe , Tongjiang , 700 m , 3.VII.2013 , coll. GQ He and LJ Xu ; Chongqing : 8 ♀ , Shaniyuan , Mt. Jinyun , 10–11.IX.2009 , coll. XC Du ; 2 ♀ , Daheba , Mt , Jinfo , 800–850 m , 15.VII.2010 , coll. XC Du and LF Song ; 1 ♂ , Baiyunzhuhai , Mt. Jinyun , 550 m , 16.VII.2011 , coll. XC Du et al. ; 1 ♂ , Laosimian , Mt. Simian , 1280 m , 14.VII.2012 , coll. GQ He et al. ; 1 ♀ , Wuli , Qianjiang , 870 m , 23.VII.2012 , coll. J Zhang and LJ Xu ; 1 ♀ , Mt. Jinyin , Qianjiang , 1100 m , 25.VII.2012 , coll. J Zhang and LJ Xu ; 2 ♀ , Mt. Jinyun , 550 m , 15.VIII.2012 , coll. XF Yang and TT Liu ; 1 ♂ , 1 ♀ , Jindianshanzhuang , Mt. Jinyun , 840 m , 14. V .2015, coll. CR Ai and D Xu ; Guizhou Province : 1 ♂ , Chishuisuoluo , 390 m , 27. V .2000, coll. YL Du , slide No. WYP05013 ; 1 ♂ , 4 ♀ , Chishuisuoluo , 390 m , 21–23.IX.2000 , coll. HL Yu ; 4 ♀ , Lanjiang , Xishui , 500 m , 24–26.IX.2000 , coll. HL Yu , slide Nos. WYP05063, WYP05064 ; 1 ♂ , Huguosi , Mt. Fanjing , 300 m , 1.VIII.2001 , coll. HH Li and XP Wang ; Yunnan Province : 23 ♂ , Mt. Weibao , Weishan , 2200 m , 20.VII.2001 , coll. HH Li and XP Wang , slide Nos. WYP05010, WYP05038, WYP05039, WYP05043, WYP05044 ; 1 ♂ , Zijiaosuo , Jingdong , 1140 m , 20. VI .2012, coll. ZG Zhang ; 1 ♂ , Mt. Ailao , Jingdong County , 2476 m , 3.VII.2013 , coll. SR Liu et al .; 20 ♂ , 6 ♀ , Mt. Weibao , Weishan County , 2244 m , 21–24.VII.2013 , coll. SR Liu et al ., slide Nos. RH15145 ♂ , RH15146 ♀ ; 25 ♂ , 5 ♀ , Mt. Weibao , Dali , 2205 m , 30.VII–2.VIII.2014 , coll. KJ Teng et al .; 1 ♂ , Malipo County , 1098 m , 4. VI .2015, coll. MF Tao ; 1 ♀ , Xiaojiangxi Village , Fengyi , Dali , 2338 m , 29.VIII.2015 , coll. KL Liu and JX Zhao ; Shaanxi Province : 3 ♂ , Xunyangba , Ningshan , 1360 m , 3.VII.2007 , HL Yu; 1 ♀ , Xunyangba , Ningshan County , 1400 m , 4.VIII.2014 , coll. HL Yu and JY Luo ; 1 ♀ , Wenjiamiao , Zhen’an , 1127 m , 28.VII.2016 , coll. JX Zhao and P Liu. Adult ( Fig. 12 ) wingspan 21.0–29.0 mm. Diagnosis. This species is characterized by the oval uncus with the long and needle-shaped lateral process arising from posterior 1/3 of lateral side, trifurcate distal process of the gnathos, sacculus with the outer dorsal process serrated on the outer margin, and median sclerotized plate almost semicircular ( Fig. 27 ). It is is similar to L. furcellata sp. nov. in the male genitalia, and differences between the two species are stated in the diagnosis of L. furcellata sp. nov. Distribution. China ( Anhui , Chongqing , Fujian , Gansu , Guangdong , Guangxi , Guizhou , Hainan , Hebei , Henan , Hubei , Hunan , Jiangsu , Jiangxi , Shaanxi , Shanxi , Sichuan , Taiwan , Tianjin , Xinjiang , Yunnan , Zhejiang ), India , Indonesia , Korea , Myanmar , Russia , Sri Lanka ( Li, Ren et al. , 2009 ).