Diversity of the genus Tmarus Simon, 1875 from Xiaolong Mountains in western China (Araneae: Thomisidae)
Author
Zhang, Rui
0000-0001-6937-6434
Key Laboratory of Zoological Systematics and Application of Hebei Province, Institute of Life Science and Green Development, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China & 15531220981 @ 163. com; https: // orcid. org / 0000 - 0001 - 6937 - 6434
Author
Zhang, Feng
0000-0002-3347-1031
Key Laboratory of Zoological Systematics and Application of Hebei Province, Institute of Life Science and Green Development, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China & dudu 06042001 @ 163. com; https: // orcid. org / 0000 - 0002 - 3347 - 1031
text
Zootaxa
2023
2023-06-08
5301
1
75
93
http://dx.doi.org/10.11646/zootaxa.5301.1.3
journal article
54602
10.11646/zootaxa.5301.1.3
c23a50a4-ed4c-4acc-8265-42b51c1a1330
1175-5326
8027829
1A5F53A2-09C3-467F-9831-A9B99071BF6D
Tmarus orientalis
Schenkel, 1963
Figs 10–17
Tmarus orientalis
Schenkel, 1963: 183
, figs 105a–b (description male).
Song & Zheng 1981: 350
, figs 5–8 (description female).
For full list of taxonomic references see
WSC (2023)
.
Material examined.
CHINA
:
Gansu Province
:
2♁
3♀
,
Qingshui County
,
Shanmen Town
,
Shanmen Village
,
34°41′24″N
,
106°21′43″E
,
1630m
,
24 June 2022
,
Zhaoyi Li
leg.
;
4♀
,
Zhang County
,
Xinsi Town
,
Heihe Forest
Farm
,
34°38′58″N
,
104°37′33″E
,
1676m
,
1 July 2022
,
Rui Zhang
leg.
Diagnosis.
Males of this species resemble those of
T. qinlingensis
Song & Wang, 1994
(
Song & Zhu 1997: 53
, figs 29 A–D) in having RTA with sharp apex, but can be distinguished from the latter by the following combination of characters: (1) embolar base slightly wider than embolar tip (vs. embolar base much wider than embolar tip in
T. qinlingensis
); (2) embolar tip bent downwards (vs. straight in
T. qinlingensis
). Females of
T. orientalis
are also similar to those of
T. qinlingensis
in having separate copulatory openings and short, thick copulatory ducts, but can be distinguished from the latter by the following characters: (1) posterior margin of atrium not obvious (vs. Vshaped in
T. qinlingensis
); (2) hood arched (vs. absence in
T. qinlingensis
).
Redescription. Male:
total length 4.19. Prosoma 1.76 long, 1.56 wide; opisthosoma 2.12 long, 1.35 wide. Carapace brown, with yellow markings. Eye sizes and interdistances: AME 0.06, ALE 0.12, PME 0.08, PLE 0.09, AME–AME 0.21, AME–ALE 0.17, PME–PME 0.25, PME–PLE 0.37, AME–PME 0.33, ALE–PLE 0.23. MOA 0.33 long, front width 0.21, back width 0.25. Leg measurements: I 8.26 (2.36, 0.82, 2.06, 1.88, 1.14); II 8.06 (2.35, 0.86, 1.98, 1.79, 1.08); III 4.08 (1.22, 0.63, 1.06, 0.58, 0.59); IV 4.43 (1.51, 0.41, 1.05, 0.77, 0.69). Leg formula: 1243. Abdomen narrow and long, posteriorly wide, dorsal greyish black, with long spines and brown spots at the base of the spines.
Palp (
Figs 14–15
). VTA digitiform, with curved apex; RTA with sharp apex. Tegulum flat; spermophor visible, around the whole tegulum, with U-shape apex. Embolus thick and strong, originating from the retrolateral flank of tegulum, about ~ 3 o’clock position.
FIGURES 10–13.
Tmarus orientalis
Schenkel, 1963
. 10–11 Male habitus (10 dorsal, 11 ventral); 12–13 Female habitus (12 dorsal, 13 ventral).
FIGURES 14–17.
Tmarus orientalis
Schenkel, 1963
. 14–15 Left male palp (14 ventral, 15 retrolateral); 16 Epigynum, ventral view; 17 Vulva, dorsal view. A = atrium; CD = copulatory duct; CO = copulatory opening; E = embolus; FD = fertilization duct; H = hood; RTA = retrolateral tibial apophysis; S = spermatheca; T = tegulum; VTA = ventral tibial apophysis.
Female:
total length 4.64. Prosoma 1.97 long, 1.79 wide; opisthosoma 2.67 long, 1.83 wide. Eye sizes and interdistances: AME 0.07, ALE 0.09, PME 0.15, PLE 0.12, AME–AME 0.21, AME–ALE 0.19, PME–PME 0.32, PME–PLE 0.43, AME–PME 0.34, ALE–PLE 0.32. MOA 0.34 long, front width 0.21, back width 0.32. Leg measurements: I 7.01 (1.93, 1.04, 1.68, 1.35, 1.01); II 7.04 (2.07, 1.06, 1.59, 1.40, 0.92); III 4.33 (1.32, 0.59, 1.09, 0.66, 0.67); IV 4.45 (1.55, 0.53, 0.95, 0.81, 0.61). Leg formula: 2143. Body color, shape and markings as in males.
Epigynum (
Figs 16–17
). Epigynum wider than long, with a transverse curved ridge; copulatory openings under the ridge; copulatory ducts thick and short; spermathecae reniform. Fertilisation ducts short, nearly as long as 1/4 spermathecal length.
Distribution.
China
(
Hebei
,
Shandong
,
Shanxi
,
Henan
,
Shaanxi
,
Gansu
),
Korea
.