Systematic revision of Gonaphodiellus taxa, with description of two new genera and fourteen new species (Coleoptera: Scarabaeidae: Aphodiinae) Author Dellacasa, Marco Museo di Storia Naturale e del Territorio Università di Pisa Via Roma, 79 I- 56011 Calci (Pisa), Italy dellacasa@museo.unipi.it Author Dellacasa, Giovanni Via Talamone 31 / 19 I- 16127 Genova, Italy dellacasag@alice.it Author Gordon, Robert D. Northern Plains Entomology P. O. Box 65 Willow City, ND 58384 USA rdgordon@utma.com text Insecta Mundi 2012 2012-04-06 2012 230 1 41 journal article 55533 10.5281/zenodo.5174222 b0a0b4e7-7c0d-4b51-9a59-156975677df7 1942-1354 5174222 Gonaphodiellus bimaculosus ( Schmidt, 1909 ) ( Fig. 6-12 ) Aphodius bimaculosus Schmidt, 1909: 19 . Aphodius ( Gonaphodiellus ) bimaculosus ; Schmidt 1913: 134 ; Dellacasa 1988: 98 . Aphodius ( Gonaphodiellus ) xalapensis Galante et al. 2003: 304 (new synonymy) . Gonaphodiellus bimaculosus ; Skelley et al. 2007: 4 . Type locality. Mexico . Type repository. Naturhistoriska Riksmuseet, Stockholm , Sweden ( type examined). Redescription. Length 5.0-6.0 mm, oblong, moderately convex, shiny, glabrous. Brown-yellowish; head, pronotum disc and elytra brownish, elytra usually with a yellowish round spot on preapical declivity ( Fig. 10 ), sometimes elytra entirely brownish ( Fig. 9 ); legs dark brown; antennal club brownish. Head with epistome feebly convex on disc, subregularly and rather densely punctured; clypeus anteriorly subtruncate, obtusely round at sides, lateral margins finely bordered, edge very shortly sparsely bristled; genae obtusely round, very shortly ciliate, protruding from the eyes; frontal suture finely impressed; front rather sparsely regularly punctured. Pronotum transverse, moderately convex, subregularly punctured; punctation somewhat denser on sides; lateral margins almost parallelsided, finely bordered, edge very shortly sparsely bristled; hind angles widely truncate, inwardly sinuate; base bisinuate. Scutellum triangularly elongate, with few sparse and irregular punctures on basal half. Elytra slightly broadened posteriorly, with a small denticle at shoulder; striae fine, superficially punctured, faintly crenulate; interstriae entirely shiny, slightly convex, finely sparsely punctured. Hind tibiae upper spur as long as half of first tarsal segment; first tarsal segment as long as following four segments combined. Male: pronotum not narrowed anteriorly, relatively somewhat more sparsely punctured on disc; fore tibiae slender and more elongate, their spur stouter, downward bent and outward curved; metasternal plate longitudinally feebly excavate at middle; aedeagus Fig. 11-12 . Female: pronotum slightly narrowed anteriorly, relatively somewhat more densely punctured on disc; fore tibiae stouter and shorter, their spur slender and almost straight; metasternal plate flat. Material examined. EL SALVADOR : Monte Cristo , 23 Km N Matapan , m 2300, 08-10.V.1971 , leg. Howden H. F. (12 exx., CMNO ) . MEXICO : Hidalgo : 2.9 Km S Molango , 20.VII.1988 . leg. Turnbow R. (2 exx., RHTC ) ; 1 mi. S Santa Monica , 6100’, 12.VII.1973 , leg. Newton A. (4 exx. CMNO , DCGI ) ; 7 Km SW Tenango de Doria , VII.1971 , leg. Newton A. (2 exx., CMNO ) ; 6.4 mi. S Tenango de Doria , m 3000, 24-28.VII.1969 , leg. Peck S. & J., cloud forest, oak (5 exx., CMNO ) ; Tlanchinol env., m 1500, 10-24.VIII.2001 , leg. Kabourek V. (2 exx., DCGI ) ; 6.4 mi. S Tianguistengo , 6800’, 12.VII.1973 . leg A. Newton ( 1 ex. , DCGI ) ; Zacualtipan , 27.V.1979 , leg. Zunino M. (2 exx., DCGI ) ; Jalisco : 10 mi W Atenquique , 6700’ 18.IX.1971 , leg. Newton A. (5 exx. CMNO , DCGI ) ; Oaxaca : 13 mi. N Nochixtlan , 7000’, 27.VI.1968 , leg. Hardy A. R. , Espinosa L. & Abrayaya J. P., Hardy Central American Expedition , ( 1 ex. , USNM ) ; Km 40 Camino Teotitlan , Hualta de Jiménez , m 2400, 09.XI.1968 , leg. Reyes P. & Cabrera M , bosque nebular (2 exx., DCGI , FSCA ) ; Km 133 carr. Tuxtepec to Oaxaca-Atepec , Llano de las Flores , 17°26’37.2"N – 96°30’13.2"W , m 2600, 14.IX.2003 , leg. Martínez I. & Suarez T. ( 1 ex. , DCGI ) ; 30 mi. Valle Nacional , 17.VIII.1973 , leg. Newton A. (2 exx., CNCI ) ; Puebla : 5 mi. W Huauchimango , 5900’, 07.VII.1970 , leg. Newton A. (3 exx., CNCI ) ; 6 mi W Teziutlan , 4-6.VIII.1960 , leg. Howden H. F. ( 1 ex. , DCGI ) ; Queretaro : 25 mi. E Landa de Matamoros , 5000’, 14-17.VII.1969 , leg. Peck S. & J., liquidambar forest (2 exx., CNCI , 1 ex. , DCGI ) ; Veracruz : Mpio. Coscomatepec , Cuiyachapa , 06.II.1999 , leg. Alvarado M. (5 exx., DCGI ) ; Cuiyachapa , carr. á Potrero Nuevo , 15.IX.1994 , leg. Arellano L. & Zunino M. (3 exx., DCGI ) ; Mpio. Coscomatepec , carr. Cuiyachapa , Pueblo Nuevo , 08.XI.1999 , leg. Dellacasa M. ( 1 ex. , DCGI ) ; El Duraznal , m 1600, 15.VII.1994 , leg. Arellano L. & Sanchez R. ( 1 ex. , DCGI ) ; Mpio. Xico , Oxtlapa , m 2100, 16.XI.1999 , leg. Dellacasa M. ( 1 ex. , DCGI ) ; carr. Xico-Tonalaco , 19°26’21"N – 97°06’10"W , m 2300, 09.XII.1999 - 09.II.2000 , leg. Montes de Oca E. & Santiago Q. (4 exx., DCGI ) ; idem, 06.VII.- 11.VIII.2000 , leg. Montes de Oca E. & Santiago Q. (13 exx., DCGI ) . Distribution. El Salvador , Mexico ( Hidalgo , Jalisco , Oaxaca , Puebla , Queretaro , Veracruz ). Bionomics. Probably all year round, coprophagous species found in several kinds of dung.