<document id="98C32EBAC75CD0B9C10D9008766CDE34" ID-DOI="10.11646/zootaxa.5575.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14746442" ID-ZooBank="521D404D-EB9D-4110-989E-A8A273C10152" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737970514015" checkinUser="plazi" docAuthor="Génier, François, Gunter, Nicole L. &amp; Saxton, Natalie A." docDate="2025" docId="038987E8FFFC6E10FF4616ECFEBAF898" docLanguage="en" docName="zootaxa.5575.1.3.pdf" docOrigin="Zootaxa 5575 (1)" docSource="https://doi.org/10.11646/zootaxa.5575.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhytomus howdeni Génier &amp; Gunter &amp; Saxton 2025, new combination" docType="treatment" docVersion="3" lastPageNumber="98" masterDocId="FFB0FF90FFF76E1BFFD11005FFC3FFB0" masterDocTitle="Rhytomus Génier &amp; Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="110" masterPageNumber="87" pageNumber="98" updateTime="1737971966986" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C6B8B13DD7102FCAE8FB435DB35B4A86">Rhytomus Génier &amp; Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="9EABE8C056C444D87CAA404CE3BAFC17">Génier, François</mods:namePart>
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<mods:namePart id="91A4A2DD7A1911DF85EDFBE973D1B4A4">Gunter, Nicole L.</mods:namePart>
<mods:affiliation id="25F3999F3DF2B919A7A94F9DE790C433">Biodiversity and Geosciences Program, Queensland Museum, South Brisbane, QLD, 4101 Australia.</mods:affiliation>
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<mods:namePart id="4BD714325FA18C58CFFCA59393703329">Saxton, Natalie A.</mods:namePart>
<mods:affiliation id="DD64AC49AD607AC453DCBCEA87CB7BB7">Department of Biology, Case Western Reserve University, 2080 Adelbert Road, Cleveland, OH, 44106, USA. &amp; Corresponding author</mods:affiliation>
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<taxonomicName id="4C204D7DFFFC6E10FF4616ECFD86F8B4" authority="(Paulian, 1985)" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="Paulian" baseAuthorityYear="1985" box="[151,581,1769,1796]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="98" phylum="Arthropoda" rank="species" species="howdeni" status="comb. nov.">
<emphasis id="B954EAECFFFC6E10FF4616ECFE43F8B4" bold="true" box="[151,384,1769,1796]" italics="true" pageId="11" pageNumber="98">Rhytomus howdeni</emphasis>
(
<bibRefCitation id="EFB14B0FFFFC6E10FE5E16EFFDFFF8B4" author="Paulian, R." box="[399,572,1770,1796]" pageId="11" pageNumber="98" pagination="219 - 238" refId="ref16422" refString="Paulian, R. (1985) Les coleopteres Scarabaeidae Canthonides de Nouvelle-Guinee. Annales de la Societe Entomologique de France, Nouvelle Serie, 21 (2), 219 - 238. https: // doi. org / 10.1080 / 21686351.1985.12278409" type="journal article" year="1985">Paulian, 1985</bibRefCitation>
)
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, 
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<figureCitation id="131B2A7BFFFC6E10FF46170BFF2EF898" box="[151,237,1806,1832]" captionStart="FIGURE 7" captionStartId="10.[152,255,1714,1738]" captionTargetBox="[156,1425,184,1681]" captionTargetId="figure-22@10.[151,1435,179,1688]" captionTargetPageId="10" captionText="FIGURE 7. Habitus of Rhytomus species in mainland New Guinea. A, R. fervidus; B, R. heurni; C, R. howdeni; D, R. larseni; E, R. latidentatus; F, R. papuanus; G, R. setulosum; H, R. sodalis; I, R. vanus; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14746458" httpUri="https://zenodo.org/record/14746458/files/figure.png" pageId="11" pageNumber="98">Fig. 7C</figureCitation>
, 
<figureCitation id="131B2A7BFFFC6E10FF28170BFEDDF898" box="[249,286,1806,1832]" captionStart="FIGURE 4" captionStartId="7.[152,255,934,958]" captionTargetBox="[151,1435,182,906]" captionTargetId="figure-17@7.[151,1435,179,909]" captionTargetPageId="7" captionText="FIGURE 4. Diagnostic characters of Rhytomus. A, dorsal head showing emargination and interocular space (R. howdeni); B, dorsal head showing lack of emargination and interocular space (R. fervidus) C, ventral view of male foreleg (R. latidentatus); D, lateral view of elytra (R. vanus); E, dorsal view of male protibia (R. latidentatus)." figureDoi="http://doi.org/10.5281/zenodo.14746450" httpUri="https://zenodo.org/record/14746450/files/figure.png" pageId="11" pageNumber="98">4A</figureCitation>
, 
<figureCitation id="131B2A7BFFFC6E10FEF8170BFE8FF898" box="[297,332,1806,1832]" captionStart="FIGURE 8" captionStartId="11.[152,255,1592,1616]" captionTargetBox="[154,1432,921,1562]" captionTargetId="figure-331@11.[151,1435,918,1565]" captionTargetPageId="11" captionText="FIGURE 8. Pygidium of Rhytomus species. A, male R. fervidus; B. female R. heurni; C, male R. howdeni; D, male R. larseni; E, male R. latidentatus; F, male R. papuanus; G, female R. setulosus; H, male R. sodalis; I, male R. vanus. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14746460" httpUri="https://zenodo.org/record/14746460/files/figure.png" pageId="11" pageNumber="98">8C</figureCitation>
, 
<figureCitation id="131B2A7BFFFC6E10FE89170BFEBAF898" box="[344,377,1806,1832]" captionStart="FIGURE 9" captionStartId="12.[152,255,1959,1983]" captionTargetBox="[155,1431,238,1929]" captionTargetId="figure-53@12.[151,1435,234,1933]" captionTargetPageId="12" captionText="FIGURE 9. Male aedeagus of Rhytomus species, left lateral, dorsal, right lateral.A, R. elongatus; B, R. fervidus; C, R. howdeni; D, R. larseni. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14746462" httpUri="https://zenodo.org/record/14746462/files/figure.png" pageId="11" pageNumber="98">9C</figureCitation>
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