A classification of Danaus butterflies (Lepidoptera: Nymphalidae) based upon data from morphology and DNA Author Smith, David A. S. Author Lushai, Gugs Author Allen, John A. text Zoological Journal of the Linnean Society 2005 2005-06-30 144 2 191 212 https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2005.00169.x journal article 10.1111/j.1096-3642.2005.00169.x 0024-4082 5434140 DANAUS GILIPPUS (CRAMER, 1775) Type locality: Brazil , Rio de Janeiro . Polytypic species (A & V-W) with some ten named subspecies ( Clark, 1941 ; Talbot, 1943 ; Urquhart, 1960 ). For range, see A & V-W, table 29.