New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae) Author Chanabun, Ratmanee Author Inkavilay, Khamla Author Panha, Somsak text ZooKeys 2017 672 1 34 http://dx.doi.org/10.3897/zookeys.672.10212 journal article http://dx.doi.org/10.3897/zookeys.672.10212 1313-2970-672-1 48810F7697B940B29A75DB0F98C5311F 48810F7697B940B29A75DB0F98C5311F Genus Glyphidrilus Horst, 1889 Diagnosis. Prostomium zygolobous. Body shape nearly circular in cross section in anterior part, and becoming quadrangular in posterior part or after clitellum. Anus dorsal or dorso-terminal. A longitudinal lamellar ridge at maturity from body wall on each side in bc, through several of the clitellar segments, which are called wings. Dorsal pores absent. Setae four pairs per segment. Clitellum annular. Genital apertures, all minute and superficial. Male pores inter-or intraclitellar. Spermathecal pores usually all behind the testis segments. Gizzard in VII or VIII sometimes extending into an adjacent segment. Calciferous glands absent. Seminal vesicles usually short, usually four pairs in IX-XII . Holonephridia. Nephrostomes single ducts avesiculate and without sphincters or caeca. Testis and funnels free in X and XI. Male ducts intramural. Ovaries fan shaped and with several egg strings. Ovisacs present or absent. Prostate glands absent and spermathecae without diverticulum ( Chanabun et al. 2013 , Gates 1972 ). Type species. Glyphidrilus weberi Horst, 1889. Type species by original designation in Horst (1889 : 77).