Order Rodentia - Family Echimyidae Author Wilson, Don E. Author Reeder, DeeAnn text 2005 The Johns Hopkins University Press Baltimore Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2 1575 1592 book chapter 0-8018-8221-4 10.5281/zenodo.7316535 Trinomys gratiosus Moojen 1948 Trinomys gratiosus Moojen 1948 , Univ. Kansas Publ. Nus Nat. Hist., 1 (19): 379 . Type Locality: Brazil , Espirito Santo , Santa Teresa, Floresta da Caixa Dagua ( 19 ^o 55’S , 40 ^o 36’W ) . Vernacular Names: Gracile Atlantic Spiny-rat . Subspecies: : Subspecies Trinomys gratiosus subsp. gratiosus Moojen 1948 Subspecies Trinomys gratiosus subsp. bonafidei Moojen 1948 Distribution: South bank of River Doce, Espírito Santo southward to Teresopolis, Rio de Janeiro ( SE Brazil ). Discussion: Recognized as a subspecies of iheringi by Moojen (1948) and Pessôa and Reis (1996). Lara and Patton (2000) elevated gratiosus to a species, retained bonafidei as a subspecies of gratiosus , and suggested that panema was not morphologically distinct from gratiosus . Karyotype has 2n=56 and FN=108 (Pessôa and Reis, 1996).