A revision of the Larainae (Coleoptera, Elmidae) of Venezuela, with description of nine new species
Author
Maier, Crystal A.
text
ZooKeys
2013
329
33
91
http://dx.doi.org/10.3897/zookeys.329.4961
journal article
http://dx.doi.org/10.3897/zookeys.329.4961
1313-2970-329-33
Nebilinagena doylei Kodada &
Jaech
, 1999
Figs 5, 54, 55, 56, 57
For complete species descriptions and genitalia illustrations, see
Kodada and
Jaech
1999
.
Material examined.
VENEZUELA:
Bolivar
State: "SE-Venezuela,
Bolivar
State; Quebrada Wiwiparu, the stream; near Paratepui village, which; crosses the tourist pathway to Mt." "Roraima, ca. 1000m a.s.l.; 31 Jan 1999; T. Derka & R. Reilmaier, lgt." (1 Holotype NMW). "VENEZUELA:
Bolivar
State;
6°5'14.1"N
,
61°23'55.8"W
400m; Piedra de la Virgen, 31.vii.2008; leg. A. Short, M. Garcia, L. Joly; AS-08-056, small forest stream" (4 specimens SEMC). "VENEZUELA:
Bolivar
State;
6°1'38"N
,
61°23'41.1"W
, 690m; Along La Escalera; 31.vii.2008; leg. A. Short, M. Garcia, L. Joly; AS-08-059; rocky stream" (4 specimens SEMC). "VENEZUELA:
Bolivar
State;
06°04'54.7"N
,
61°23'52.7"W
, 509 m; Along La Escalera, Highway 10; 14.vii.2010; leg. Short, Tellez, & Arias; VZ10-0714-01C" (2 specimens SEMC - frozen tissue collection) "VENEZUELA:
Bolivar
; Municipio Gran Sabana; El Pauji. 25/IV/2004 al 02/V/2004 J. Camacho; J. Perozo, Col." "
04°28'66"N
,
61°35'38"W
; 880msnm." (9 specimens MALUZ). "Loc. VEN-16-2010, canyon; of the Rio Yuruen above; Uruyen village, Venezuela; Bolivar Province" "NP Canaima,
5°42'26.0"N
;
62°28'13.7"W
, alt. 658m; a.s.l. 16.i.2010; leg. T. Derka & M. Svitok (11 specimens CKB).
Diagnosis.
Neblinagena doylei
can be distinguished from
Neblinagena prima
by its range, which does not overlap with the range of the latter. Additionally, in
Neblinagena doylei
the pronotal longitudinal carinae are entire and the oblique furrow of the pronotum forms a 30° angle relative to the median longitudinal furrow (Fig. 57), whereas in
Neblinagena prima
the pronotal lateral carinae are obscured anteriorly and the oblique furrows form a 45° angle relative to the median longitudinal furrow (Fig. 61).
Neblinagena doylei
is also slightly larger than its congener.
Intraspecific variation.
This species varies slightly in size (5.5-6.6 mm) and in coloration - from dark brown to black.
Distribution and habitat.
Neblinagena doylei
can be found throughout the eastern part of Venezuela, in and around the Gran Sabana, in
Bolivar
and Amazonas States (Fig. 5). Specimens have been collected on flat, waterslide-like rocks in the La Escalera region.
Associated species.
This species is associated the Mount Roraima endemic species
Roraima carinata
. Other aquatic beetles collected at the same localities include:
Notionotus
spp.,
Oocyclus
spp. (
Hydrophilidae
), and
Macrelmis
spp. (
Elmidae
:
Elminae
).