Annotated type catalogue of the Bulimulidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London
Author
Breure, Abraham S. H.
Naturalis Biodiversity Center, P. O. Box 9517, Leiden, the Netherlands
bbreure@xs4all.nl
Author
Ablett, Jonathan D.
Natural History Museum, Division of Higher Invertebrates, London, SW 7 5 BD, UK
text
ZooKeys
2014
2014-03-21
392
1
367
http://dx.doi.org/10.3897/zookeys.392.6328
journal article
http://dx.doi.org/10.3897/zookeys.392.6328
1313-2970-392-1
FFCF5A59FFB1FF93FFF10B43FFAFFFF5
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Helix thamnoica
d'Orbigny
, 1835
Figs 68A
, L60ii
Helix thamnoica
d'Orbigny
1835
: 16.
Bulimus thamnoicus
d'Orbigny
1837 [1834-1847]
: 290, pl. 37 figs 4-9 [5 June 1837; text 6 May 1838].
Scutalus thamnoicus
;
Breure 1975b
: 1144, pl. 4 fig. 1.
Type locality.
"Var. A. [Not given]. Var. B. [Not given]. Var. C. Cavari (republica Boliviana). Var. D. provincia Chuquisacensi (republica Boliviana)"; see remarks and
Breure 1973
: 123.
Label.
"Var. A. Palca
d'ayupaya
Bolivia", "Var. B. Cochabamba, Bolivia", "Var. C. Cavari, Bolivia", "Var. D chuquisaca Bolivia", all in
d'Orbigny's
handwriting.
Dimensions.
"Longit. a 45 ad 80 millim.; latit. a 22 ad 39 millim.", corrected in
d'Orbigny
1837 [1834-1847]
to: "Long. ex 25 ad 65 millim.; lat. ex. 15 ad 35 millim."; figured specimen herein H 44.0, D 25.3, W 6.8.
Type material.
NHMUK 1854.12.4.111, five paralectotypes "var. A."; 1854.12.4.109, six paralectotypes "var. B."; 1854.12.4.107, four paralectotypes [var. B]; 1854.12.4.110, nine syntypes "var. C."; 1854.12.4.108, lectotype (
design. n.
) and four paralectotypes "var. D." (all material
d'Orbigny
coll.).
Remarks.
d'Orbigny's
varieties have proven to belong to different species, already noticed by
Pilsbry (1897 [1897-1898])
and
Richardson (1995)
, although they disagreed on the specific assignments of the four varieties.
Breure (1975b)
mentioned only material pertaining to the varieties A and B in the Paris museum; however, also material corresponding to variety D is available in this collection (Breure, unpublished data). The data presented in
d'Orbigny
1837-1838 [1834-1847]
, together with the material of both the NHMUK and MNHN collections, are used here to resolve this complicated issue.
Var. A. "
Magna
,
reticulata
" (pl. 37 figs 4-5) is recorded from
"pres
de Palca, capitale de la province
d'Ayupaya"
[Dept. Cochabamba, Indepencia; see
Breure 1973
: 133, fig. 6] and "Longueur, 45
millimetres"
. The lectotype (MNHN 23258) has a shell height of 47.7 mm; the largest specimen in NHMUK labelled "var. A" has shell height 44 mm. The name
thamnoica
d'Orbigny
is restricted to this variety.
Var. B. "
Magna
,
crassa
,
striata
" (pl. 37 fig. 7) is listed from "environs de
Capinata
[Dept. La Paz]" and from
"vallee
de Cochabamba (...) principalement
a
Viloma. Longueur, 65
millimetres"
; see
Breure 1973
: 133, figs 6-7. The largest specimen in lot NHMUK 1854.12.4.109 measures 52.7 mm in height (
Fig. 68B
); note that there is also material without variety designation (1854.12.4.107), of which the largest specimen has a shell height of 61.5 mm. This variety is considered to be identical with
Bulimus alauda
Hupe
, 1857.
Var. C. "
Minor
,
fasciata
" (pl. 37 fig. 8) has been found at "les coteaux du bourg de Cavari", near the top of the mountains; see
Breure 1973
: 129, fig. 7. "Longueur, 25
millimetres"
. The largest specimen in NHMUK has dimensions H 32.2, D 19.0, W 6.3. This variety was considered a synonym of
Bulimus revinctus
Hupe
, 1857 by both
Pilsbry (1897 [1897-1898])
and
Richardson (1995)
. We consider this variety as
Bulimus gayi
Pfeiffer, 1857.
Var. D. "
Marmorata
" (pl. 37 fig. 9) was collected "aux environs de la ville de Chuquisaca"; see
Breure 1973
: 130, fig. 4. No shell height was given by
d'Orbigny
. The largest NHMUK specimen, corresponding to
d'Orbigny's
figure, has dimensions H 40.8, D 23.1, W 6.2 (
Fig. 68C
). Three specimens are present in Paris (MNHN 23261), of which the largest one with dimensions H 45.3, D 23.9, W 6.7.
Pilsbry (1897 [1897-1898])
regarded var. D as synonymous with
Bulimus alauda
Hupe
, 1857;
Richardson (1995)
synonymyzed it with
Bulimus revinctus
Hupe
, 1857. The types of both species have not been located, leaving only the possibility to check against
Hupe's
figures (1857: pl. 7 fig. 3 resp. pl. 7 fig. 2), in which case we concur with
Pilsbry's
view.
Current systematic position.
Bulimulidae
,
Kuschelenia (Kuschelenia)
species. See under remarks.