Contribution to the knowledge of the genus Dendarus Dejean, 1821 (Coleoptera: Tenebrionidae: Blaptinae: Dendarini) from Iran, Turkmenistan and some adjacent territories
Author
Nabozhenko, Maxim V.
Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str., 45, Makhachkala, Republic of Dagestan 367000, Russia. & Dagestan State University, M. Gadzhiev str., 43 a, Makhachkala, Republic of Dagestan 367000, Russia.
Author
Poggi, Roberto
Museo Civico di Storia Naturale “ Giacomo Doria ”, Genova, 16121 Italy.
text
Zootaxa
2022
2022-06-20
5155
1
105
123
journal article
77865
10.11646/zootaxa.5155.1.5
008f04ee-4fad-4629-b397-6aae0a9db700
1175-5326
6669239
7964FEBD-3A6A-4E68-B867-C1214D49F491
Dendarus vagabundus vagabundus
Reitter, 1904
, sp. resurr.
(
Figs 7
,
8
,
9G
,
10F
,
11F
,
12F
,
13G
)
Reitter, 1904: 84
.
=
D. vagans
Reitter, 1902: 180
(junior homonym, nec
D. vagans
Seidlitz, 1893: 399
);
=
Dendarus armeniacus
sensu
Medvedev, 1968: 98
, 108, figs 106–108;
Medvedev & Nepesova, 1985: 123
;
Medvedev & Nepesova, 1990
a
: 25
;
Medvedev & Nepesova, 1990
b
: 886
.
=
D. transcaspicus transcaspicus
sensu
Kaszab, 1973: 53
, 54, fig. 8;
=
D. transcaspicus medvedevi
Kaszab, 1973: 55
, fig. 7, photo 16,
syn. n.
Type material.
Holotype
of
Dendarus vagans
,
♂
(
HNHM
): “Transcasp. Kisil-Aswat” [now
Turkmenistan
:
Serdar
], “
transcaspicus
Branczik
det. dr. Kaszab”, “
Dendarus vagans
i.l. m 1899”, “Typus
Dendarus vagans
Reitt. Coll. Reitter
”.
Material.
Turkmenistan
.
3♂♂
,
1♀
(
ZIN
):
Suchudly
,
vii.1901
(leg.
C. Saaro
)
;
1♂
(
ZIN
):
Kopetdag
, Kel’te-
Chinar
,
30.viii.1935
(leg.
K. Arnoldi
)
;
1♂
,
1♀
(
ZIN
):
Dushak
, 20 from
Firuza
,
21.ix.1935
(leg.
K. Arnoldi
)
;
1♀
(
ZIN
):
Kopetdag
,
20 km
from
Firyuza
[now Archabil’],
Chaekh
spring,
24.ix.1935
(leg.
K. Arnoldi
)
;
1♂
,
1♀
(
ZIN
):
Shakh-Shakh Mt.
,
13.x.1935
(leg.
K. Arnoldi
)
;
1♂
(
ZIN
):
Kopetdag
,
Tamil’
canyon,
Chandyr River valley
,
29.x.1935
(leg.
K. Arnoldi
)
;
1♀
(
ZIN
):
W Kopetdag
,
Ay-Dere
[canyon],
8.iv.1952
(leg.
K. Borisova
)
;
1♂
(
ZIN
):
W Kopetdag
,
Ay-Dere
,
29–30.iv.1952
(leg.
Shteinberg
)
;
1♂
,
1♀
(
ZIN
):
Ay-Dere
,
0–3 km
from river mouth,
30.iv.1952
(leg.
Romadina
)
;
1♂
,
1♀
(
ZIN
):
Ay-Dere
, forestry,
30.iv.1952
(leg.
Odintsova
)
;
1♀
(
ZIN
):
Kyzyl-Arvat
[now Serdar],
3.v.1952
(leg.
Il’ichev
)
;
2♂♂
(
ZIN
):
W Kopetdag
,
Syunt Mt.
,
1800 m
,
14.v.1952
(leg.
Osipova
)
;
2♂♂
,
2♀♀
(
ZIN
):
W Kopetdag
,
Syunt Mt.
,
14.v.1953
(leg.
Shteinberg
)
;
1♂
,
1♀
(
ZIN
):
W Kopetdag
,
Syunt Mt.
,
21.vi.1953
(
O.L. Kryzhanovskij
)
;
4♀♀
(
ZIN
):
Kara-Kala
[now Garrygala], N slope of
Syunt Mt.
,
6.iii.1956
(leg.
G.S. Medvedev
)
;
2♂♂
,
5♀♀
(
ZIN
):
Kara-Kala
, canyon on N slope of
Syunt Mt.
,
6.v.1957
(leg.
G.S. Medvedev
)
;
6♂♂
,
5♀♀
(
ZIN
):
Kara-Kala
,
Savudzhe
canyon,
8.v.1957
(leg.
G.S. Medvedev
)
;
1♂
,
1♀
(
ZIN
):
Kara-Kala
,
Khasar
,
2.10.1961
(
L. Freiberg
)
;
4♂♂
,
3♀♀
(
ZIN
):
Ay-Dere
,
28–29.iv.1971
(leg.
G.S. Medvedev
)
;
17♂♂
,
16♀♀
(
ZIN
): N slope of
Syunt Mt.
,
28.iv.1974
,
2.v.1974
,
4.v.1974
(leg.
G.S. Medvedev
)
;
2♂♂
,
5♀♀
(
ZIN
): N slopes of
Syunt
and
Khasardag Mts
,
30.iv.1974
(leg.
G.S. Medvedev
)
;
1♂
,
1♀
(
ZIN
):
Kopetdag
,
Ay-Dere
,
21.iv.1976
(leg. D. Yanushev)
;
1♀
(
ZIN
):
Kopetdag
,
10 km
E Ipaykala
guard,
20.v.1984
(leg.
G.S. Medvedev
)
;
4♂♂
,
4♀♀
(
ZIN
):
6 km
SO
Terskan
,
14.v.1984
(leg.
G.S. Medvedev
)
;
1♂
with
Cyrillic
label (
HNHM
):
Kazandzhik
env., mountains,
600 m
,
21.iv.1952
(leg.
O.L. Kryzhanovskij
)
;
1♀
with
Cyrillic
label (
HNHM
): footnotes
18 km
S Kyzyl-Arvat
[now Serdar],
22.iv.1952
(leg.
Il’ichev
)
;
1♂
(
HNHM
):
Kopet-Dagh Mts.
,
Dushak
,
37°57ʹN
,
57°54ʹE
,
2200 m
, 1–
2.10.1991
,
No L
33 (leg.
Podlussány, L. Ronkay, Z. Varga
)
.
Iran
.
2♂♂
(
ZIN
):
Imam-Gulyar
,
Kuchan
[now
Razavi Khorasan Province
, Quchan],
16.iii.1898
(leg.
N. Zarudny
)
;
1♀
(
MSNG
):
Khor Vall
[e]y [now
East Azerbaijan Prov.
:
Khor Khor Darrehsî
],
Oct
[ober] [19]73 (unknown collector)
;
1♂
(
HNHM
):
Mazandaran
/
Tehran
provinces, between
Polour
and
Abali
,
2100–2600 m
,
17.v.1970
(leg.
Wittmer
,
v. Bothmer
)
;
2♂♂
(
HNHM
):
Mazandaran Prov.
,
Abharm
(
Damavend
),
2100–2400 m
,
15.v.1970
(leg.
Wittmer
,
v. Bothmer
)
.
Diagnosis.
Male. Body slender, from moderately (
Fig. 8D, E
) to very shiny (
Fig. 8A
), legs and antennae slender, long. Head with moderately coarse, sparse puncturation (puncture diameter subequal to interpuncture distance). Head ventrally with simple (not impressed) longitudinal furrow beyond prementum. Pronotum moderately convex. Lateral margins of pronotum evenly rounded, emarginated or rarely straight at basal quarter; anterior angles not protruding or rarely slightly protruding; base bisinuate, posterior angles rectangular, weakly protruding. Pronotum punctured by uniform fine and sparse round punctures. Prosternal process comparatively wide, rounded at apex. Elytra widest at middle, with lateral margins less rounded. Strial punctures moderate in size or large, densely spaced (35–40 punctures in one row), round, not impressed or slightly impressed. Interstriae with fine and sparse distinctly visible puncturation. Protibiae narrow, with deep and wide notches on inner side and protruding, weakly obtuse tooth before the indentation; protibia widest at level of tooth. Mesotibiae usually straight, sometimes weakly bent. Mesotarsomeres 1–4 not widened, tarsomere 2 with reduced hair brush on sole, very rarely mesotarsomere 3 also with very small reduced hair tuft.
FIGURE 7.
Dendarus vagabundus
, male, holotype of
D. vagans
(junior homonym). A = habitus, laterally; B = habitus, dorsally; C = abdomen; D = dissected aedeagus and tergite VII; E = labels.
FIGURE 8.
D. vagabundus
, habitus. A = male, dorsally (Iran, North Khorasan Province: Bojnurd); B = male, ventrally (Turkmenistan, Kopetdag); C = female, dorsally (Turkmenistan, Kopetdag); D = male, dorsally (Iran, Golestan Prov.: Khoshyeylag); F = female, dorsally (the same locality).
FIGURE 9.
Dendarus
spp.
, male protibia. A =
D. armeniacus
; B =
D. crenulatus
; C =
D. leonhardi
; D =
D. simplex
; E =
D. matthewsi
sp. n.
; F =
D. transcaspicus
; G =
D. vagabundus
.
FIGURE 10.
Dendarus
spp.
, male tergite VIII. A =
D. armeniacus
; B =
D. crenulatus
; C =
D. leonhardi
; D =
D. matthewsi
sp. n.
; E =
D. transcaspicus
; F =
D. vagabundus
.
Female. Body much more robust, protibiae weakly bent. Lateral margins of pronotum always widely emarginated in basal quarter; posterior angles acute, stronger protruding.
Notes.
Dendarus vagans
Reitter, 1902
and
D. vagabundus
Reitter, 1904
were considered by
Kaszab (1973)
as junior synonyms of
D. transcaspicus transcaspicus
Brancsik, 1899
as a result of misidentification.
The
holotype
of
D. vagabundus medvedevi
Kaszab, 1973
(studied by the first author in HNHM but not photographed) from Saramsakli (now
Turkmenistan
, Central Kopetdag, Sarymsakly spring,
38°29ʹ11ʺ N
,
57°6ʹ49ʺ E
) is only a deviant wrinkled specimen of
D. vagabundus vagabundus
with a less protruding tooth on the inner side. So, we propose the following synonymy:
Dendarus vagabundus vagabundus
Reitter, 1904
=
D. transcaspicus medvedevi
Kaszab, 1973
,
syn. n.
Dendarus vagabundus afghanicus
Kaszab, 1973
from Central
Afghanistan
(Sang Mazar, Pur Chaman) is a clear subspecies, which differs from the nominotypical one by the deeper notch and right tooth on inner side of male protibia and denser and coarser puncturation of the pronotum.
Distribution.
Iran
(recorded for
Iran
by
Medvedev & Nepesova (1990
a
)
under the name
D. armeniacus
),
Turkmenistan
.