Resurrection of Pareas yunnanensis (Vogt, 1922) with description of a new species of Pareas from Yunnan Province, China (Squamata, Pareidae) Author Liu, Shuo 84A37AC1-E7EA-4E47-9BE4-65EC4C96B6C4 Kunming Natural History Museum of Zoology, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan 650223, China. & Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan 650201, China. liushuo@mail.kiz.ac.cn Author Zhang, Dongru E8F2FC4E-069F-4657-95F3-1356C1A73A42 Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan 650201, China. zhangdongru@mail.kiz.ac.cn Author Poyarkov, Nikolay A. 39C4DC5F-374F-486E-9DFB-22420286CEE6 Faculty of Biology, Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory 1 / 12, Moscow 119234, Russia. & Laboratory of Tropical Ecology, Joint Russian-Vietnamese Tropical Research and Technological Center, Nghia Do, Cau Giay, Hanoi, Vietnam. n.poyarkov@gmail.com Author Hou, Mian 26774FFD-255C-40E6-ABFD-8021DC2B6714&E27890C7-34C5-4546-91E6-BE097AF9C257 College of Continuing (Online) Education, Sichuan Normal University, Chengdu, Sichuan 610066, China. & Yunnan Academy of Forestry and Grassland, Kunming Yunnan 650201, China. turtlechina@126.com&125701986@qq.com Author Wu, Li E27890C7-34C5-4546-91E6-BE097AF9C257 Yunnan Academy of Forestry and Grassland, Kunming Yunnan 650201, China. 125701986@qq.com Author Rao, Dingqi AEB67455-CBA4-418B-9EBF-626F7DAD5299 Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, Yunnan 650201, China. raodq@mail.kiz.ac.cn Author Nguyen, Tan Van B13846C0-32AC-4EEC-A9E5-C28F910CA62C Institute for Research and Training in Medicine, Biology and Pharmacy, Duy Tan University, Da Nang, 550000, Vietnam. tan.sifasv@gmail.com Author Vogel, Gernot 26C84E11-E43F-4076-B268-28C32E9FD2D0 Society for Southeast Asian Herpetology, Im Sand 3, Heidelberg D- 69115, Germany. gernot.vogel@t-online.de text European Journal of Taxonomy 2023 2023-02-15 860 1 1 26 http://dx.doi.org/10.5852/ejt.2023.860.2045 journal article 55044 10.5852/ejt.2023.860.2045 5177db24-e558-4aea-bf3e-4ae656d6c216 2118-9773 7651005 D489D635-D4C1-46A3-B535-7882386EFF06 Pareas ( Eberhardtia ) yunnanensis ( Vogt, 1922 ) Figs 2–4 ; Tables 3–4 Amblycephalus yunnanensis Vogt, 1922: 142 . Amblycephalus yunnanensis — Mell 1922 . — Pope 1935 . — Ding et al . 2020 . Amblycephalus monticola yunnanensis — Mell 1931 . Pareas yunnanensis — Rao & Yang 1992 . — Guo et al . 2020 — Wang et al . 2020 . — Liu & Rao 2021 . Pareas chinensis (in part) — Zhao & Adler 1993 . — Zhao 2006 . — Yang & Rao 2008 . — Wallach et al . 2014 . Pareas cf. yunnanensis — Poyarkov et al . 2022 . Revised diagnosis This species differs from Pareas geminatus by having fewer subcaudals, relatively more rows of keeled mid-dorsal scales, relatively fewer ventral scales, solid black dorsal surface of head, and no distinct large black spots or stripe on each side of head; it differs from P. xuelinensis Liu & Rao, 2021 by infralabials not fused with chin-shields, vertebral scales enlarged, having fewer ventral scales, fewer subcaudals, relatively more rows of keeled mid-dorsal scales, and solid black dorsal surface of head. SVL 387–482 mm in adults, TL 94–110 mm in adults, TL/SVL 0.22–0.25; preoculars mostly single, rarely two; postoculars and suboculars mostly fused, rarely separated; loreal bordering orbit in most individuals; prefrontal bordering orbit; supralabials 6–8, infralabials 6–8; infralabials not fused with chin-shields; dorsal scales in 15 rows throughout the body; vertebral scales enlarged; 5–7 rows of middorsal scales keeled on the middle part of the body; precloacal plate undivided; ventral scales 169–175; subcaudals 59–65, all paired. Dorsal surface of head black; no or 1–2 indistinct large black spots on each side of head, no stripe on each side of head; two wide black stripes pass from parietals to the vertical black bars on neck; dark nuchal band absent; vertical black bars distinct on trunk and indistinct on tail; iris brownish-yellow or brownish-orange. Etymology The specific epithet is a toponymic adjective given in reference to Yunnan Province , China . We propose “ Yunnan slug-eating snake” for the common English name and “ṙ南钝±蛇” (Yún Nán Dùn Tóu Shé) for the common Chinese name of this species. Type material Syntypes CHINA • 2 ♀♀ (adults); Dali Prefecture , Yunnan Province ; ZMB 27660 , 65431 . Additional material examined CHINA – Yunnan Province • 2 ♀♀ adults; Dali Prefecture , Dali City , Xiaguan Town ; 25°36′17″ N , 100°11′33″ E ; elev. 2250 m ; 12 Jun. 2022 ; S. Liu leg.; KIZ 2022033 , 2022034 • 1 ♀ adult; Dali Prefecture , Dali City , Dali Town ; 25°41′56″ N , 100°8′2″ E ; elev. 2230 m ; 13 Jun. 2022 ; S. Liu leg.; KIZ 2022035 • 1 juv. ; Dali Prefecture , Dali City , Xiaguan Town ; 25°37′52″ N , 100°11′44″ E ; elev. 2140 m ; 15 Jun. 2022 ; S. Liu leg.; KIZ 2022036 . Fig. 2. Pareas yunnanensis ( Vogt, 1922 ) , syntypes in preservative. A–D . ZMB 27660. A . Dorsal view. B . Ventral view. C–D . Close up views of the head. E–H . ZMB 65431. E . Dorsal view. F . Ventral view. G–H . Close up views of the head. Photos by G. Vogel. Table 3. Morphological characters of Pareas yunnanensis ( Vogt, 1922 ) . For abbreviations, see Material and methods section. Measurements in mm, paired meristic characters given as left/right, “?” = not available, data in parentheses from Vogt (1922) .

ZMB 27660 Syntype	ZMB 65431 Syntype	KIZ 2022033	KIZ 2022034	KIZ 2022035	KIZ 2022036
SEX	♀	♀	♀	♀	♀	Juvenile
SVL	387 (385)	405 (410)	482	395	445	163
TL	? (95)	? (100)	105	94	110	39
TL/SVL	? (0.25)	? (0.24)	0.22	0.24	0.25	0.24
PrFBO	Yes/Yes	Yes/Yes	Yes/Yes	Yes/Yes	Yes/Yes	Yes/Yes
PreO	1/1	1/1	1/1	1/1	2/1	1/1
PosO	Fused	Fused	Fused	Fused	2/2	Fused
SubO	Fused	Fused	Fused	Fused	1/1	Fused
SPOF	Yes	Yes	Yes	Yes	No	Yes
ATem	2/2	2/2	1/1	1/1	2/2	1/1
PTem	3/2	3/3	2/2	2/2	2/3	2/2
SupL	?	7/6	7/8	7/7	7/7	6/6
InfL	?	7/7	7/8	8/6	8/8	7/6
LoBO	Yes/Yes	No/Yes	Yes/No	No/No	Yes/Yes	Yes/Yes
Vs	172	175	173	174	169	173
Prec	Single	Single	Single	Single	Single	Single
Sc	65	64	59	60	64	61
Ds	15-15-15	15-15-15	15-15-15	15-15-15	15-15-15	15-15-15
NED	1	1	1	1	1	1
NKD	7	5	7	7	7	5
Max	?	?	4/4	5/5	4/4	5/4
DNB	Absent	Absent	Absent	Absent	Absent	Absent

Distribution This species is currently known only from Dali City and its adjacent areas, Dali Prefecture, Yunnan Province , China ( Fig. 8 ). Fig. 3. Pareas yunnanensis ( Vogt, 1922 ) , specimens from its type locality in preservative. A . Dorsal views. B . Ventral views. Photos by S. Liu. Fig. 4. Pareas yunnanensis ( Vogt, 1922 ) , specimens from its type locality in life. A–C . ♀, KIZ 2022033. D–F . ♀, KIZ 2022034. G–I . ♀, KIZ 2022035. Photos by S. Liu. Table 4. Comparisons of morphometric and scalation data for Pareas yunnanensis ( Vogt, 1922 ) and P. chinensis (Barbour, 1912) . Measurements in mm, data for P. chinensis (restricted to Sichuan Province, China) from Poyarkov et al . (2022) .

Pareas yunnanensis	Pareas chinensis
SVL	385–482	309–427
TL	95–110	99–128
TL/SVL	0.22–0.25	0.26–0.33
PrFBO	Yes	Yes
PreO	1–2	1–2
PosO	Fused or 2	Fused or 1
SubO	Fused or 1	Fused or 1
SPOF	Yes or No	Yes or No
ATem	1–2	2
PTem	2–3	3–4
SupL	6–8	6–8
InfL	6–8	7–9
LoBO	Yes or No	No
Vs	169–175	169–178
Sc	59–65	69–76
Ds	15-15-15	15-15-15
NED	1	0 or 3
NKD	5–7	0 or 5
Max	4–5	5–6
DNB	Absent	Absent

Phylogenetic position Pareas yunnanensis belongs to the subgenus Eberhardtia and is a member of the P. hamptoni ( Boulenger, 1905 ) species group ( Poyarkov et al . 2022 ). Phylogenetically it is reconstructed as a sister species of P. geminatus (see Fig. 1 ).