A new species of Cephalocyclus (Coleoptera: Scarabaeidae: Aphodiinae) from Mexico Author Minor, Pablo Author Dellacasa, Marco Author Dellacasa, Giovanni text Zootaxa 2015 3974 2 297 300 journal article 10.11646/zootaxa.3974.2.13 bb2bcafd-f29b-4d82-bd43-e74e89f1db34 1175-5326 246067 C21E4DBC-C745-4890-A78A-87A035DABC03 Cephalocyclus moroni Minor, Dellacasa , & Dellacasa, new species ( Figs. 1–4 ) Type locality. Las Joyas, Sierra de Manantlán, Autlán, Jalisco, Mexico . Holotype repository. Instituto de Ecología, A. C. collection ( IEXA , Xalapa, Mexico ). Description. Holotype . Male : Length from clypeus to pygidium 7.5 mm ; humeral width 3.0 mm. Oval, elongate, moderately convex, shiny. More or less dark brown; clypeal margin and sides of pronotum red; antennal club brown; legs dark brown with tarsi paler, shiny. Head with epistoma feebly convex on disc; finely, somewhat irregularly punctate; clypeus faintly sinuate at middle, round at sides, rather thinly bordered, edge glabrous. Genae angulate, sparsely ciliate, distinctly protruding from the eyes. Frontal suture finely impressed; front distinctly, unevenly punctate. Pronotum transverse, moderately convex, somewhat flattened at sides mainly toward front and hind angles; evenly, finely, and sparsely punctate; lateral margins feebly arcuate, thickly bordered, edge subcerciniform, glabrous; hind angles subtruncate; base not bordered. Scutellum triangular, microreticulate, sparsely punctate. Elytra oval elongate, moderately convex, feebly widened posteriorly; striae fine, superficially punctate, subcrenulate; interstriae faintly convex, superficially microreticulate, sparsely and finely punctate. Elytra pubescent laterally and on preapical declivity only, humerus dentate, epipleura distinct and sparsely ciliate. Metatibia with upper spur longer than first tarsomere; first tarsomere as long as the following three tarsomeres combined. Protibial spur relatively elongate and stout; metasternal plate concave; aedeagus ( Fig. 4 ). TABLE 1. Checklist of the genus Cephalocyclus Dellacasa, Dellacasa , & Gordon, 1998 (type species: Aphodius fuliginosus Harold, 1863 by original designation). Species Distribution 1 Cephalocyclus bordati Dellacasa, Dellacasa, & Gordon, 2007 Mexico (Chihuahua, Durango) 2 Cephalocyclus carmenae Dellacasa, Dellacasa, & Gordon, 2007 Mexico (Coahuila, Nuevo León) 3 Cephalocyclus cartagoensis Dellacasa, Dellacasa, & Gordon, 2007 Costa Rica 4 Cephalocyclus costaricensis Dellacasa, Dellacasa, & Gordon, 2007 Costa Rica

5	Cephalocyclus durangoensis (Bates, 1887) [ Aphodius ] Synonym: Aphodius hogei durangoensis Bates, 1887 Synonym : Aphodius pugil Balthasar, 1946	Mexico (Coahuila, Durango, Michoacán, Nuevo León, Oaxaca)
6	Cephalocyclus fuliginosus (Harold, 1863) [ Aphodius ]	Mexico (Chihuahua, Distrito Federal, Durango, Guanajuato, Hidalgo, Jalisco, México, Michoacán, Morelos, Oaxaca, Puebla, Sinaloa, Tlaxcala, Zacatecas)
7	Cephalocyclus gravidus (Harold, 1863) [ Aphodius ]	Mexico (Chihuahua, Distrito Federal, Durango, Hidalgo, Jalisco, México, Michoacán)

8 Cephalocyclus halffteri Dellacasa, Dellacasa, & Gordon, 2007 Mexico (Guerrero, México ) 9 Cephalocyclus hogei (Bates, 1887) [ Aphodius ] Mexico (Durango, Hidalgo, México , Michoacán, Synonym: Aphodius transversus Robinson, 1940 Morelos, Oaxaca, Puebla, Querétaro, Veracruz); United States of America (Arizona?) 10 Cephalocyclus howdenorum Dellacasa, Dellacasa, & Gordon, 2007 Mexico (Nuevo León)

11	Cephalocyclus lagoi Dellacasa, Dellacasa , & Gordon, 2007	Mexico (Coahuila, San Luis Potosí)
12	Cephalocyclus luridiventris (Harold, 1862) [ Aphodius ]	Mexico (Distrito Federal, Oaxaca, Puebla)
13	Cephalocyclus luteolus (Horn, 1887) [ Aphodius ]	United States of America (Arizona, New Mexico, Texas, Utah)

14 Cephalocyclus majomaensis Dellacasa, Dellacasa, & Gordon, 2013 Mexico (Zacatecas) 15 Cephalocyclus mexicanus (Harold, 1862) [ Aphodius ] Mexico (Distrito Federal, Durango, Guerrero, Synonym: Aphodius omiltemius Bates, 1890 Hidalgo, Jalisco, México , Michoacán, Morelos, Nayarit, Oaxaca, Puebla, Querétaro, Sinaloa, Veracruz) 16 Cephalocyclus moroni Minor, Dellacasa , & Dellacasa, 2015 Mexico (Jalisco) 17 Cephalocyclus ordonezi Dellacasa, Dellacasa, & Gordon, 2007 Mexico ( México ) 18 Cephalocyclus potosinus Dellacasa, Dellacasa, & Gordon, 2007 Mexico (Nuevo León) 19 Cephalocyclus pseudofuliginosus Dellacasa, Dellacasa , & Gordon, Mexico (Oaxaca) 2008 20 Cephalocyclus pullatus (Schmidt, 1913) [ Aphodius ] Mexico (Durango,Sinaloa) 21 Cephalocyclus puncticeps (Harold, 1862) [ Aphodius ] Mexico (Jalisco, México , Puebla) 22 Cephalocyclus rockefelleri Dellacasa, Dellacasa, & Gordon, 2007 Mexico (Chihuahua) 23 Cephalocyclus stebnickae Deloya & Ibañez-Bernal, 2000 Mexico (Chiapas) 24 Cephalocyclus villosipes (Harold, 1862) [ Aphodius ] Mexico (Hidalgo, México , Oaxaca, Puebla, Veracruz) FIGURES 1–4. Cephalocyclus moroni new species (Las Joyas, Sierra de Manantlán, Autlán, Jalisco, Mexico): 1–2. habitus of male; 3. epipharynx; 4. aedeagus (lateral view). Allotype . Similar to holotype except as follows: Length from clypeus to pygidium 7.0 mm; humeral length 2.8 mm , head and pronotum less transverse and more convex, protibial spur elongate and slender; metasternal plate flat. Variation . Males vary in length from 7.0–8.0 mm, humeral length from 2.8–3.0 mm. The head and pronotum more transverse and less convex than females. One male present the head and pronotum reddish brown. Females vary in length from 7.0– 7.5 mm , humeral length from 2.8–3.0 mm. Type material. MEXICO : Jalisco, Autlán, S. Manantlán, Las Joyas, light Hg, 01-VIII-2002 . 1,900 m . E. García- Real, J. Allende, A. Ramírez y M. A. Morón cols. Eight specimens ( holotype male, allotype female, three male and three female paratypes ) at IEXA ; two specimens (one male and one female paratypes ) at MXAL ; three specimens (one male and two female paratypes ) at DCGI . Distribution. Known from type locality only. Natural history. Specimens of the type series were collected in August and were attracted to light. Etymology. Named in honor of Dr. Miguel Angel Morón, Mexican specialist of the Scarabaeoidea of Mexico and Latin America . Discussion. Cephalocyclus moroni is most similar to Cephalocyclus ordonezi Dellacasa, Dellacasa, & Gordon, 2007 , because both species share the dentate elytral humerus, but C. moroni is longer and wider, the pronotum and elytral interstices are sparsely pubescent, and both sexes have protibial with apical spur longer than the first tarsomere. The males of C. ordonezi have the apical spur abruptly bent downward and outwardly curved. Cephalocyclus ordonezi is known only from the mountains of Puebla and Estado de México , nearly 580 km south of Sierra de Manantlán, in localities between 3,000 to 3,800 m of elevation; whereas C. moroni was collected at 1,900 m of altitude. Furthermore, some character states of C. moroni are also shared with Cephalocyclus stebnickae Deloya & Ibañez-Bernal, 2000 , which is only known from San Cristóbal de las Casas, Chiapas, more than 1,200 km south of Sierra de Manantlán, both species have the pronotal and clypeal margins not ciliate.