<document ID-DOI="10.11646/zootaxa.5255.1.33" ID-ISSN="1175-5326" ID-Zenodo-Dep="7745110" ID-ZooBank="8D7A551D-646D-49E2-A9AA-A14EACC67777" checkinTime="1679053428171" checkinUser="plazi" docAuthor="Misirlioğlu, Mete, Reynolds, John Warren, Stojanović, Mirjana, Trakić, Tanja, Sekulić, Jovana, James, Samuel W., Csuzdi, Csaba, Decaëns, Thibaud, Lapied, Emmanuel, Phillips, Helen R. P., Cameron, Erin K. &amp; Brown, George G." docDate="2023" docId="7D2487ECFFB71B73FF3EFE3EFE04FD66" docLanguage="en" docName="zootaxa.5255.1.33.pdf" docOrigin="Zootaxa 5255 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5255.1.33" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tumakidae Righi 1995" docType="treatment" docVersion="3" lastPageNumber="429" masterDocId="811DFF94FFBB1B7FFFA9FFB6FFF4FFA5" masterDocTitle="Earthworms (Clitellata, Megadrili) of the world: an updated checklist of valid species and families, with notes on their distribution" masterLastPageNumber="438" masterPageNumber="417" pageNumber="429" updateTime="1679083476692" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title>Earthworms (Clitellata, Megadrili) of the world: an updated checklist of valid species and families, with notes on their distribution</mods:title>
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<mods:namePart>Misirlioğlu, Mete</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-9928-8478</mods:nameIdentifier>
<mods:affiliation>Eskisehir Osmangazi University, Faculty of Science and Letters, Department of Biology, TR- 26040 Eskisehir, Turkey. https: // orcid. org / 0000 - 0001 - 9928 - 8478</mods:affiliation>
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<mods:namePart>Reynolds, John Warren</mods:namePart>
<mods:affiliation>Oligochaetology Laboratory, 9 - 1250 Weber Street East, Kitchener, ON Canada N 2 A 4 E 1, and Research Associate, New Brunswick Museum, Saint John, NB, Canada E 2 K 1 E 5.</mods:affiliation>
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<mods:namePart>Stojanović, Mirjana</mods:namePart>
<mods:affiliation>University of Kragujevac, Faculty of Science, Department of Biology and Ecology, Radoja Domanovića 12, 34000 Kragujevac, Serbia.</mods:affiliation>
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<mods:namePart>Trakić, Tanja</mods:namePart>
<mods:affiliation>University of Kragujevac, Faculty of Science, Department of Biology and Ecology, Radoja Domanovića 12, 34000 Kragujevac, Serbia.</mods:affiliation>
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<mods:namePart>Sekulić, Jovana</mods:namePart>
<mods:affiliation>University of Kragujevac, Institute for Information Technologies Kragujevac, Department of Sciences, Jovana Cvijića bb, 34000 Kragujevac, Serbia.</mods:affiliation>
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<mods:namePart>James, Samuel W.</mods:namePart>
<mods:affiliation>Maharishi International University, Department of Regenerative Agriculture, Fairfield, IA, 52557, USA.</mods:affiliation>
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<mods:namePart>Csuzdi, Csaba</mods:namePart>
<mods:affiliation>Dept. of Zoology, Eszterházy Károly Catholic University, Eger, Hungary.</mods:affiliation>
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<mods:namePart>Decaëns, Thibaud</mods:namePart>
<mods:affiliation>CEFE, Univ Montpellier, CNRS, EPHE, IRD, Montpellier, France.</mods:affiliation>
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<mods:namePart>Lapied, Emmanuel</mods:namePart>
<mods:affiliation>Taxonomia Biodiversity Fund, 7 rue Beccaria, 72012, Paris, France.</mods:affiliation>
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<mods:namePart>Phillips, Helen R. P.</mods:namePart>
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<mods:namePart>Cameron, Erin K.</mods:namePart>
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<mods:namePart>Brown, George G.</mods:namePart>
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<mods:date>2023</mods:date>
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<taxonomicName ID-CoL="84KWP" authority="Righi, 1995" authorityName="Righi" authorityYear="1995" box="[151,441,392,418]" class="Clitellata" family="Tumakidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="429" phylum="Annelida" rank="family">
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Tumakidae 
<bibRefCitation author="Righi, G." box="[299,441,392,418]" pageId="12" pageNumber="429" pagination="485 - 607" refId="ref19527" refString="Righi, G. (1995) Colombian earthworms. In: Van der Hammen, T. &amp; Santos, A. G. (Eds.), Studies on Tropical Andean Ecosystems, vol. 4. Cramer, Berlin, pp. 485 - 607." type="book chapter" year="1995">Righi, 1995</bibRefCitation>
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<paragraph blockId="12.[151,1437,464,707]" pageId="12" pageNumber="429">
The monogeneric 
<taxonomicName authorityName="Righi" authorityYear="1995" box="[357,482,464,490]" class="Clitellata" family="Tumakidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="429" phylum="Annelida" rank="family">Tumakidae</taxonomicName>
family has only three known species, 
<taxonomicName authority="Righi" authorityName="Righi" authorityYear="1995" box="[919,1175,464,490]" class="Clitellata" family="Tumakidae" genus="Tumak" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="429" phylum="Annelida" rank="species" species="hammeni">
<emphasis box="[919,1105,464,490]" italics="true" pageId="12" pageNumber="429">Tumak hammeni</emphasis>
Righi
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, 
<taxonomicName authority="Celis" authorityName="Celis" box="[1191,1407,464,490]" class="Clitellata" family="Tumakidae" genus="Tumak" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="429" phylum="Annelida" rank="species" species="amari">
<emphasis box="[1191,1341,464,490]" italics="true" pageId="12" pageNumber="429">Tumak amari</emphasis>
Celis
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&amp; Rangel-Ch., and 
<taxonomicName authority="Celis" authorityName="Celis" box="[341,602,500,526]" class="Clitellata" family="Tumakidae" genus="Tumak" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="429" phylum="Annelida" rank="species" species="congorum">
<emphasis box="[341,537,500,526]" italics="true" pageId="12" pageNumber="429">Tumak congorum</emphasis>
Celis
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&amp; Rangel-Ch., found in relatively drier, lower-elevation forests in Central 
<collectingCountry box="[151,261,536,562]" name="Colombia" pageId="12" pageNumber="429">Colombia</collectingCountry>
and the Caribbean region, along the 
<collectingRegion box="[691,815,536,562]" country="Colombia" name="Magdalena" pageId="12" pageNumber="429">Magdalena</collectingRegion>
River valley (
<bibRefCitation author="Righi, G." box="[982,1114,536,563]" pageId="12" pageNumber="429" pagination="485 - 607" refId="ref19527" refString="Righi, G. (1995) Colombian earthworms. In: Van der Hammen, T. &amp; Santos, A. G. (Eds.), Studies on Tropical Andean Ecosystems, vol. 4. Cramer, Berlin, pp. 485 - 607." type="book chapter" year="1995">Righi 1995</bibRefCitation>
; Celis &amp; Rangel-Ch. 2015). Given the relatively small amount of remaining forest fragments of its native habitat in 
<collectingCountry box="[1131,1244,572,598]" name="Colombia" pageId="12" pageNumber="429">Colombia</collectingCountry>
, due primarily to agricultural expansion and pastures for cattle, further efforts are needed to ensure adequate conservation of the few species known. Additional sampling efforts are also needed to discover if there are any other species present along its potential distribution range.
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