Late Devonian-early Carboniferous ostracods (Crustacea) from South China: taxonomy, diversity and implications Author Guillam, Elvis 66D41887-865C-4B91-861F-ADB09B801962 Centre de Recherche en Paléontologie - Paris, MNHN - SU - CNRS; 8 rue Buffon, 75005 Paris, France. elvis.guillam@sorbonne-universite.fr Author Forel, Marie-Béatrice 063C1F7E-6D26-48F9-A8B1-2AFA496B5FB8 Centre de Recherche en Paléontologie - Paris, MNHN - SU - CNRS; 8 rue Buffon, 75005 Paris, France. marie-beatrice.forel@mnhn.fr Author Song, Junjun 5676B30E-AF63-45DE-A465-2EC8BF85ABF9 Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Palaeoenvironment, Chinese Academy of Sciences, Nanjing, China. jjsong@nigpas.ac.cn Author Crasquin, Sylvie 6FDDD550-8522-404D-B3A2-D30BFAA86927 Centre de Recherche en Paléontologie - Paris, MNHN - SU - CNRS; 8 rue Buffon, 75005 Paris, France. sylvie.crasquin@sorbonne-universite.fr text European Journal of Taxonomy 2022 2022-03-15 804 1 62 http://dx.doi.org/10.5852/ejt.2022.804.1689 journal article 55868 10.5852/ejt.2022.804.1689 987c26cd-d47c-4773-8b5a-df53f0eda18e 2118-9773 6359315 5EEDA63F-8A81-40DF-B5A4-C799498DC523 Kloedenellitina spinosa Gurevitch, 1972 Fig. 6B–C Kloedenellitina spinosa Gurevitch, 1972: 333, pl. 11 figs 3–4. Kloedenellitina spinosa – Tschigova 1977: 171 , pl. 41 fig. 13. Material examined CHINA • 1 complete carapace (♀, Fig. 6B ); Blue Snake section , Gelaohe Formation , sample 19BAI 60; P6M 3857 • 1 complete carapace (♂, Fig. 6C ); Blue Snake section , Gelaohe Formation , sample 19BAI 69; P6M 3858 • 16 complete carapaces ; Blue Snake section , Gelaohe Formation , samples 19BAI 60, 19BAI 67–69, 19BAI 74, 19BAI 80. All from the Famennian , late Devonian . Dimensions RV: L = 552–783 µm, H = 307–394µm, H/L = 0.48–0.59. LV: L = 434–761 µm, H = 210–377 µm, H/L = 0.45–0.56. Remarks In the genus Kloedenellitina , females have an inflated posterior carapace, which corresponds to a brood pouch ( Ellis & Messina 1964 ). This character allows us to distinguish females ( Fig. 6B ) from males ( Fig. 6C ) in the studied material. The lateral surface of the specimens preserving their carapace appears as reticulate ( Fig. 6C ), it is rather smooth in steinkern but the reticulation remains visible in certain areas ( Fig. 6B ). Our material is smaller than the type material (L = 760–930 µm, H = 440–490 µm in Gurevitch 1972) with H/L between 0.53 and 0.58 while it is between 0.45 and 0.59 in our material (both valves taken together). The specimen shown in Tschigova (1977 : pl. 61 fig. 13) is also larger with L = 1200 mm , H = 556mm but the H/L ratio is similar (H/L = 0.46). In the present material, this species may mainly be represented by juvenile specimens corresponding to at least two different ontogenetic stages that we cannot discriminate because of the wide distribution of their dimensions. Table 2 (continued on next two pages). Stratigraphic distribution of species in the Blue Snake section.

Orange: Association A, Grey: Association B, Green: Association C.
Gelaohe Formation					Tangbagou Formation
Famennian			latest Fa.			Tournaisian
Sample 19BAI Species	80	74	69	68	67	63	60	58	55	54	44	43	41	39	37	27-2	23	16	08	05	03	02	01
Knoxina sp.	1	8
Bairdia nanbiancunensis	1	2	1	2
Fellerites sp. 1	2	1	1	1	3
Beyrichiopsis sp.	8	1	2	3	1
Kloedenellitina spinosa	1	2	2	2	3	1	7
Kloedenellitina sygmaeformis	3	2	1	6	3	3
Knoxiella subcompressa	2	1	2	2	1
Fellerites sp. 2	2	2	5	1	1	2
Knoxiella complanata	2	1	4	6	2	9	17	5
Bairdia hypsela	2	6	1	1	7	1	2	14	2	2	1	9	7	16	3	6
Parabolbinella sp.	1
Sulcella baisuzhena sp. nov.	1	1	19
Clavofabella ? lanshella sp. nov.	2	4	17	1	1
Kloedenellitina sincera	14	1	2	3
Sansabella gelaohensis sp. nov.	73	8	2	3	15	5
Healdianella sp.	1	12	3	7
Acratia acutiangulata	5	15	1	2	4	2
Bairdia quasikuznecovae	1	4	1	2	7	3	1	1
Acratia cf. tschudovoensis	1	1	8	1
Bairdiacypris nanbiancunensis	1	1	1	2	3	1	1	2	2
Clavofabella ? sp.	1	14
Selebratina sp.	1	7	2
Cytherellina caerulea sp. nov.	5	4	2	1
Healdianella cf. insolita	1	1	1	1	1
Hypotetragona sp.	13	12	8	1	3	1	1	1	1
Bairdia cf. susoroides	2	1	2	1	1
Knoxiella cf. subcompressa	2
Cavellina cf. dushanensis	2	1
Bairdia semichatovae	1	6	1	1
Bairdia ( R .) angulatiformis	1	1	3	1	1	1	1

Table 2 (continued). Stratigraphic distribution of species in the Blue Snake section.

Microcheilinella cf. decora	8	1	1	2	1	1	1
Praepilatina adamszaki	1	1	1	2	4	2	2
Bairdia obliqua	2	1	2	4	3	6	1	4	1	3	9	2	2	12	10
Paraparchites sp. 2	1
Cavellina cf. recta	2
Bairdiocypris marginifera	1
Fabalicypris sundarijanata	4	2
Acratia cf. evlanensis	1	1	1	3
Knoxiella archedensis	1	1	2	1	4	2	2	8	1	2
Elliptocyprites sp.	1	2	1	1
Bairdia sp. 3	2	2	2
Bairdioidea ? indet	1	1
Acratia subobtusa	2	1	2
Elliptocyprites lorangeri	1
Bairdia cf. extenuata Bairdia ( C .) curta McCoy, 1844 emend. Jones & Kirkby, 1878 in Crasquin, 1986	1 1
Healdianella faseollina	2	1
Bairdia sp. 5	5	1	1	1	1	2
Healdianella alba	1	1	3	2	3	3	2	9	9	9
Microcheilinella larionovae	1	1	1	1	1	3	1
Microcheilinella sp.	4	1	4	3	1	1	3
Microcheilinella sp. sensu Olempska, 1979	1	1	1	3
Bairdia ( R .) sp. cf. R . sinuosa (Morey, 1936) in Crasquin, 1985	1	5	1	2	1	2
Healdianella lumbiformis	16	2	1	1
Bairdiacypris subcyliпdrica	33	12	7	1	1	1	2
Shishaella hastierensis	8	2	6	5	5	7	1	7	1	3	2	1
Bairdia cestriensis	1	1
Shishaella porrecta	2	1
Bairdia submongoliensis	1	1
Bairdia ( C .) laveinei	1	1	1	1	1
Acratia cf. insolita	1	1	1	1	1	2	1	2
Acutiangulata acutiangulata	1	1	3	3	3	2
Bairdia mnemonica	4	2	4	3	2	1
Bairdia sp.1	2	1
Bairdia natiformis	1	1	2	2	1	1	2
Bairdiacypris quasielongata	1	1	1	1	1	3	3	5
Amphissites sp.	2	1	2	5	1

Table 2 (continued). Stratigraphic distribution of species in the Blue Snake section.

Blessites feluyensis	8	1	8	1
Bairdia sp. 6	1	1		1
Hypotetragona angulata	3	4	2	2	1	2	3	7	8	5
Paraparchites longmenshanensis	22	1	2	1	1	1
Bairdiocypris fomikhaensis	1	2	1	1	1	1	2	2	5
Bairdianella cuspis	1	1	2	1	3	5	7	6
Bairdia sp. 2	2
Bairdia solita	1	4	1
Microcheilinella sp. cf. M . buschminae Olempska, 1982 in Song & Gong, 2017	1	1	3
Coryellina grammi	1	4	1	1	2	1
Acratia similaris	1	2	4	2	4	3
Acratia sp.	1	3	3	1	1	1
Bairdia cf. altifrons	1	1	2
Bairdia confragosa	3	5	3	2	1	3	10	2	6
Bairdia feliumgibba	2	6	5	2	7
Microcheilinella sp. A , aff. buschminae Olempska, 1981 in Casier et al., 2004	1	1	2
Paraparchites sp. 1	1
Bairdia ( R .) buschminaae	2	2
Bairdia ( R .) cf. plebeja	14	1	1	3	2
Bairdia sp. 4	1	1	4	6	1
Bairdia sp. 7	1	1
Healdianella cf. subdistincta	2	5	2	1
Bairdia cf. grahamensis	1
Bairdia rustica	26
Bairdiocypris elliptica	3	2
Aparchites sp.	5	2	1	1
Acratia cf. disjuncta	1	3
Bairdiocypris sp.	1	2	1	2
Microcheilinella subrhomboidalis	1
Microcheilinella infradominica in Song & Gong, 2019	2	2
Knoxites cf. argulata	1	1

Occurrence Torchinsk Formation,Volyn region, Russian platform,Tournaisian, early Carboniferous(Gurevitch 1972). Lebedyan horizon, Torchinsk Formation, Central regions of the Russian platform, Famennian, late Devonian ( Tschigova 1977 ). Samples 19BAI 60, 19BAI 67–69, 19BAI 74, 19BAI 80, Gelaohe Formation, Blue Snake section, Famennian, late Devonian (this work).