An investigation into the taxonomy of Abavorana luctuosa (Peters, 1871) (Anura, Ranidae) and the resurrection of Rana decorata Mocquard, 1890 from Borneo Author Quah, Evan S. H. Institute of Tropical Biodiversity and Sustainable Development, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia, https: // orcid. org / 0000 - 0002 - 5357 - 1953 & Herpetology Laboratory, Department of Biology, La Sierra University, Riverside, California, USA, https: // orcid. org / 0000 - 0001 - 8422 - 3698 & Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, 117377 Singapore, https: // orcid. org / 0000 - 0002 - 0638 - 9198 Author Grismer, L. Lee Herpetology Laboratory, Department of Biology, La Sierra University, Riverside, California, USA, https: // orcid. org / 0000 - 0001 - 8422 - 3698 Author Wood Jr., Perry L. https://orcid.org/0000-0003-3767-5274 Department of Biological Sciences & Museum of Natural History, Auburn University, Auburn, Alabama 36849, USA, https: // orcid. org / 0000 - 0003 - 3767 - 5274 Author Lim, Kelvin K. P. Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, 117377 Singapore, https: // orcid. org / 0000 - 0002 - 0638 - 9198 Author Imbun, Paul Y. Museum of Zoology, Research & Education Division, Sabah Parks, 10626 Kota Kinabalu, Sabah, Malaysia Author Anuar, M. S. Shahrul School of Biological Sciences, Universiti Sains Malaysia, 11800 Minden, Penang, Malaysia, https: // orcid. org / 0000 - 0003 - 0648 - 7318 & Center for Marine and Coastal Studies, Universiti Sains Malaysia, 11800 USM, Pulau Pinang, Malaysia text Vertebrate Zoology 2021 2021-03-11 71 75 99 http://dx.doi.org/10.3897/vz.71.e60921 journal article http://dx.doi.org/10.3897/vz.71.e60921 2625-8498-71-75 1C61F5FE1FA44083B91313FD42157FCA C87DE9A5FDFA5B5FA5603914E35A1446 Abavorana luctuosa (Peters, 1871) Common Mahogany Frog Figs 3 , 4 , 7E-H , 8A-F , 9A-B Limnodytes luctuosus : Peters 1871 :579, 1872:43 & pl. 6; Capocaccia 1957 :208 & 215. Rana luctuosa (in part): Boulenger 1882 :68, 1891:341, 1912:238, 1920:183 & 184; Flower 1896 :904, 1899:896 & 897; Hanitsch 1900 :73; Butler 1904 :199 & 200; Van Kampen 1923 :196 & 197; Smith 1930 :94 & 103, 1931:16; Capocaccia 1957 : 215; Taylor & Elbel 1958:1040; Taylor 1962 :451-453; Inger 1954 :250, 1966:206-208, 1978:312, 1985 :iii, 5, 38, 40, 56, 57, 76; Grandison 1972 :66 & 67; Berry 1975 :74 & 75; Dring 1979 :200; Inger & Stuebing 1992:42, 2005:19, 27, 33, 149 & 150; Inger & Tan 1996:563; Inger et al. 1996 :363; Manthey & Grossmann 1997:111 & 112; Khonsue & Thirakhupt 2001:73; Malkmus et al. 2002 : 163-165; Das 2006a :5; Jaafar et al. 2008 :39. Rana (Limnodytes) luctuosa : Mocquard 1890 :122. Rana (Hylorana) luctuosa (in part): Boulenger 1920 :126. Rana (Hylarana) luctuosa (in part): Van Kampen 1923 :196 & 197; Bourret 1942 :360; Dubois, 1987: 42. Hylorana luctuosa (in part): Deckert 1938 :144. Rana (Pulchrana) luctuosa (in part): Dubois 1992 :326. Pulchrana luctuosa (in part): Frost et al. 2006 :369; Fei et al. 2010 :33. Hylarana luctuosa (in part): Manthey 1983 :22; Che et al. 2007 : 1-13 (by implication); Das et al. 2007 :159, 160, 167, 170, fig. 7d; Imbun 2014 :99. Abavorana luctuosa : Oliver et al. 2015 :186 (in part); Quah et al. 2017 : 272-288 (in part); Zainudin et al. 2017 :876-891 (in part); Chan et al. 2019 :1057, 1062. Hylarana (Abavorana) luctuosa (in part): Inger et al. 2017 :147 & 148. Holotype. Adult female (MSNG 29344) collected by Giacomo Doria and Odoardo Beccari from "Sarawak" , (Borneo) Malaysia ( Capocaccia 1957 ) (Fig. 3 ). Diagnosis. Body robust, medium-sized; head moderate; snout short, rounded, canthus rostralis smoothly rounded; interorbital space broader than the upper eyelid; tympanum distinct, not quite two-thirds the size of the eye with no pale colouration on the margins of the tympanum; no vocal sacs in males; vomerine teeth in two small oblique patches on a level with the posterior edge of the choanae; length of 1st finger greater than 2nd finger; disc width to finger width ratios of finger 3 and toe 4 is 1-1.5; dorsolateral fold indistinct or absent; the humeral gland in males is prominent, raised and centrally positioned on the ventral surface of the upper arm; a weak or absent rictal ridge; outer metatarsal tubercle weak or absent; subarticular tubercles moderate; skin of dorsum smooth or finely shagreened; throat, abdomen, and flanks smooth; posterior section of venter and back of the thigh rugose; dorsum reddish-orange to chocolate-brown, encircled by a white or cream coloured dorsolateral line that encircles the snout, canthus rostralis, outer edge of the upper eyelids, and dorsum along the dorsolateral fold to the vent; lower flanks dark-brown or black below the dorsolateral line grading into a paler venter; dorsal colouration of the limbs same as the flanks with whitish or light-grey speckles or stripes. Abavorana luctuosa can be easily differentiated from its congeners on the basis of its ventral colour pattern which is usually immaculate or with only very faint, sparse, light speckling (Figs 3 , 4 , 9A-B ). Adult males with SVL 35.05-44.27 mm, adult females with SVL 37.05-51.06 mm; adult males with SW 7.11-8.62 mm, adult females with SW 7.65-9.38 mm; adult males with IOD 3.25-5.14 mm, adult females with IOD 3.80-5.64 mm; adult males with TD 2.50-3.83 mm, adult females with TD 2.95-4.24 mm; adult males with FL 17.81-22.04 mm, adult females with FL 19.13-26.37 mm (Table 5 ). Distribution. Its distribution spans southern Peninsular Thailand, Peninsular Malaysia, Borneo, and Sumatra (Manthey & Grossmann 1997; Oliver et al. 2015 ; Quah et al. 2017 ). Within Borneo it is confirmed from Sabah state: Imbak Canyon and Sipitang; Sarawak state: Bukit Kana, Samarakan and Tubau in Bintulu Division and Kubah National Park where it is sympatric with Abavorana decorata (Fig. 10 ). The species is expected to range across Borneo.