<document ID-DOI="http://dx.doi.org/10.3897/BDJ.8.e52779" ID-PMC="PMC7314865" ID-Pensoft-Pub="1314-2828-8-e52779" ID-Pensoft-UUID="C17C76D8DE8352FBB21CB34B1B6DC36D" ID-PubMed="32595414" ModsDocID="1314-2828-8-e52779" checkinTime="1592451974453" checkinUser="pensoft" docAuthor="Pham, Anh Van, Ziegler, Thomas &amp; Nguyen, Truong Quang" docDate="2020" docId="EE02685510345FCB9417352EA84D5C56" docLanguage="en" docName="BiodivDatJour 8: e52779" docOrigin="Biodiversity Data Journal 8" docSource="http://dx.doi.org/10.3897/BDJ.8.e52779" docTitle="Protobothrops cornutus" docType="treatment" docVersion="2" id="C17C76D8DE8352FBB21CB34B1B6DC36D" lastPageNumber="52779" masterDocId="C17C76D8DE8352FBB21CB34B1B6DC36D" masterDocTitle="New records and an updated checklist of snakes from Son La Province, Vietnam" masterLastPageNumber="52779" masterPageNumber="52779" pageNumber="52779" updateTime="1668128426429" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>New records and an updated checklist of snakes from Son La Province, Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pham, Anh Van</mods:namePart>
<mods:affiliation>Tay Bac University, Quyet Tam Ward, Son La City, Son La Province, Vietnam</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ziegler, Thomas</mods:namePart>
<mods:affiliation>AG Zoologischer Garten Koeln, Riehler Strasse 173, D- 50735, Cologne, Germany &amp; Institute of Zoology, University of Cologne, Zuelpicher Street 47 b, D- 50674, Cologne, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nguyen, Truong Quang</mods:namePart>
<mods:affiliation>Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Hanoi, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-6601-0880</mods:nameIdentifier>
<mods:nameIdentifier type="email">nqt2@yahoo.com</mods:nameIdentifier>
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<mods:titleInfo>
<mods:title>Biodiversity Data Journal</mods:title>
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<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>8</mods:number>
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<mods:extent unit="page">
<mods:start>52779</mods:start>
<mods:end>52779</mods:end>
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<mods:url>http://dx.doi.org/10.3897/BDJ.8.e52779</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.8.e52779</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2828-8-e52779</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:EE02685510345FCB9417352EA84D5C56" httpUri="http://treatment.plazi.org/id/EE02685510345FCB9417352EA84D5C56" lastPageNumber="52779" pageId="0" pageNumber="52779">
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<taxonomicName LSID="EE026855-1034-5FCB-9417-352EA84D5C56" authority="(Smith, 1930)" baseAuthorityName="Smith" baseAuthorityYear="1930" class="Reptilia" family="Viperidae" genus="Protobothrops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protobothrops cornutus" order="Squamata" pageId="0" pageNumber="52779" phylum="Chordata" rank="species" species="cornutus">Protobothrops cornutus (Smith, 1930)</taxonomicName>
</paragraph>
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<subSubSection pageId="0" pageNumber="52779" type="materials_examined">
<paragraph pageId="0" pageNumber="52779">Materials</paragraph>
<paragraph pageId="0" pageNumber="52779">
<materialsCitation collectingDate="2015-08-26" collectionCode="Reptilia" country="Vietnam" latitude="21.210001" location="near Pap Village" longLatPrecision="21" longitude="104.76611" pageId="0" pageNumber="52779" specimenCode="TBU. MD. 2015.110" specimenCount="1" specimenCount-male="1" typeStatus="Other material">
<emphasis bold="true" pageId="0" pageNumber="52779">Type status:</emphasis>
<typeStatus pageId="0" pageNumber="52779">Other material</typeStatus>
. 
<emphasis bold="true" pageId="0" pageNumber="52779">Occurrence:</emphasis>
catalogNumber: 
<specimenCode pageId="0" pageNumber="52779">TBU. MD.2015.110</specimenCode>
; individualCount: 
<specimenCount pageId="0" pageNumber="52779">1</specimenCount>
; sex: 
<specimenType pageId="0" pageNumber="52779">male</specimenType>
; lifeStage: 
<specimenType pageId="0" pageNumber="52779">adult</specimenType>
; 
<emphasis bold="true" pageId="0" pageNumber="52779">Taxon:</emphasis>
scientificNameID: Protobothropscornutus; scientificName: Protobothropscornutus; class: Reptilia; order: Squamata; family: Viperidae; genus: Protobothrops; specificEpithet: cornutus; scientificNameAuthorship: (Smith, 1930); 
<emphasis bold="true" pageId="0" pageNumber="52779">Location:</emphasis>
country: 
<collectingCountry pageId="0" pageNumber="52779">Vietnam</collectingCountry>
; countryCode: VN; stateProvince: Son La; county: Phu Yen; municipality: Muong Do; locality: 
<location LSID="urn:lsid:plazi:treatment:EE02685510345FCB9417352EA84D5C56:CA0442DC741F3475C16DFC14D55D9889" country="Vietnam" latitude="21.210001" longLatPrecision="21" longitude="104.76611" name="near Pap Village" pageId="0" pageNumber="52779">near Pap Village</location>
; verbatimElevation: 
<elevation pageId="0" pageNumber="52779">720 m</elevation>
; verbatimLatitude: 
<geoCoordinate degrees="21" direction="north" minutes="12" orientation="latitude" precision="15" seconds="36" value="21.210001">21°12'36''N</geoCoordinate>
; verbatimLongitude: 
<geoCoordinate degrees="104" direction="east" minutes="45" orientation="longitude" precision="15" seconds="58" value="104.76611">104°45'58''E</geoCoordinate>
; verbatimCoordinateSystem: WGS84; 
<emphasis bold="true" pageId="0" pageNumber="52779">Event:</emphasis>
eventDate: 
<collectingDate pageId="0" pageNumber="52779" value="2015-08-26">26 August 2015</collectingDate>
; eventRemarks: collected by H.V. Tu and C.A.K.P Kham; 
<emphasis bold="true" pageId="0" pageNumber="52779">Record Level:</emphasis>
language: en; collectionCode: 
<collectionCode pageId="0" pageNumber="52779">Reptilia</collectionCode>
; basisOfRecord: PreservedSpecimen
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<subSubSection pageId="0" pageNumber="52779" type="description">
<paragraph pageId="0" pageNumber="52779">Description</paragraph>
<paragraph pageId="0" pageNumber="52779">
Morphological characters of the voucher specimen from Son La Province agreed well with the description of 
<bibRefCitation author="Herrmann, H W" journalOrPublisher="Herpetologica" pageId="0" pageNumber="52779" pagination="211 - 221" refId="B5672238" refString="Herrmann, H W, Ziegler, T, Malhotra, A, Thorpe, R S, Parkinson, C, 2004. Redescription and systematics of Trimeresurus cornutus (Serpentes: Viperidae) based on morphology and molecular data. Herpetologica 60: 211 - 221" title="Redescription and systematics of Trimeresurus cornutus (Serpentes: Viperidae) based on morphology and molecular data" volume="60" year="2004">Herrmann et al. (2004)</bibRefCitation>
and 
<bibRefCitation author="Luu, V Q" journalOrPublisher="Herpetology Notes" pageId="0" pageNumber="52779" pagination="149 - 152" refId="B5672270" refString="Luu, V Q, Nguyen, T Q, Lehmann, T, Bonkowski, M, Ziegler, T, 2015. New records of the horned pitviper, Protobothrops cornutus (Smith, 1930) (Sepentes: Viperidae), from Vietnam with comments on morphological variation. Herpetology Notes 8: 149 - 152" title="New records of the horned pitviper, Protobothrops cornutus (Smith, 1930) (Sepentes: Viperidae), from Vietnam with comments on morphological variation" volume="8" year="2015">Luu et al. (2015)</bibRefCitation>
: SVL 610 mm, TaL 146 mm (n = 1); body elongated; head triangular, distinct from neck, flat, covered by small scales; snout obtuse; rostral scale triangular, not visible above; nasal single, subrectangular; loreals 3/3, between nasal and preocular scale; loreal pit bordered by second supralabial and two preocular scales; supralabials 10/10, first completely separated from nasal, third supralabial largest, concave in the middle; horns including one enlarged scale and two relatively-enlarged supraoculars; supralabials separated from suboculars by 2 scales; infralabials 13/12, the first pair in contact with each other, the first three pairs in contact with the chin shields; dorsal scale rows 21-21-17, keeled; ventral scales 190; subcaudal scales 71, all paired; cloacal entire.
</paragraph>
<paragraph pageId="0" pageNumber="52779">
Colouration in life: dorsal head grey with dark brown marks; one stripe each side from posterior orbit to edge of the labial; dorsal surface grey-brown with serial blotches along the midbody; dorsolateral side light brown; ventral surface greyish-blue (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5672585" captionText="Figure 5. Protobothrops cornutus (TBU MD. 2015.110, adult male) from Son La Province, Vietnam. Photo: H. V. Tu" figureDoi="10.3897/BDJ.8.e52779.figure5" httpUri="https://binary.pensoft.net/fig/395548" pageId="0" pageNumber="52779">5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="52779" type="distribution">
<paragraph pageId="0" pageNumber="52779">Distribution</paragraph>
<paragraph pageId="0" pageNumber="52779">
In Vietnam, this species has been reported from Lai Chau and Lao Cai provinces in the north, southwards to Quang Binh and Thua Thien Hue provinces (
<bibRefCitation author="Nguyen, S V" journalOrPublisher="Edition Chimaira, Frankfurt am Main" pageId="0" pageNumber="52779" refId="B5672320" refString="Nguyen, S V, Ho, C T, Nguyen, T Q, 2009. Herpetofauna of Vietnam. Edition Chimaira, Frankfurt am Main" title="Herpetofauna of Vietnam" year="2009">Nguyen et al. 2009</bibRefCitation>
). Elsewhere, the species has been reported from China (
<bibRefCitation author="Nguyen, S V" journalOrPublisher="Edition Chimaira, Frankfurt am Main" pageId="0" pageNumber="52779" refId="B5672320" refString="Nguyen, S V, Ho, C T, Nguyen, T Q, 2009. Herpetofauna of Vietnam. Edition Chimaira, Frankfurt am Main" title="Herpetofauna of Vietnam" year="2009">Nguyen et al. 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="52779" type="ecology">
<paragraph pageId="0" pageNumber="52779">Ecology</paragraph>
<paragraph pageId="0" pageNumber="52779">The voucher specimen was found at 20:35 h on the ground. The surrounding habitat was secondary forest composed of medium and small hardwoods, lianes and shrub. Air temperature was 25-30°C and relative humidity was 70-80%.</paragraph>
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