A revision and phylogenetic analysis of the spider genus Coptoprepes Simon (Araneae: Anyphaenidae, Amaurobioidinae) Author Werenkraut, Victoria Author Ramírez, Martín J. text Zootaxa 2009 2212 1 40 journal article 10.5281/zenodo.275145 344a109d-a561-4054-97b7-e1850acc321f 1175-5326 275145 Coptoprepes bellavista new species Figs. 8–10 , 22 b, 23d–f, 24d–f Type material: Holotype : male from Chile , Región IX (Araucanía), Cautín province, Bellavista , N shore Lago Villarrica, ca. 39º S, 72º W, 310 m , site 655, window trap, Valdivian rainforest, 15–30 December 1982 , A. Newton & M. Thayer ( AMNH ; temporary preparations male: VIW-00002, VIW-00003, female: MJR- 0 1337, VIW-00004). Paratype : female, same data as holotype . Etymology: The specific name is a noun in apposition taken from the type locality. Diagnosis: Females can be distinguished by their epigyne having an anterior trapezoidal sclerotized area, leaving the posterior margin of the epigyne mostly unsclerotized ( Fig. 10 c). Males are similar to several other Coptoprepes by having a RTA curved, longer than the tibia ( Fig. 10 c), but can be distinguished from them by the truncate tip of the primary conductor and from the similar C. contulmo n. sp. by having a ventrally concave secondary conductor ( Figs. 10 b–f, 23-d–f). Description: Male ( holotype ): Prosoma length 2.07, width 1.55, narrower in front than in the female. Ttibia/metatarsus length: I, 1.43/1.33; II, 1.20/1.32; III, 0.92/1.17; IV, 1.28/1.70. Chelicerae slightly smaller than those of female, with six (right) or seven (left) teeth on retromargin, four on promargin. Sternum length 1.13, width 0.83. Spines: leg I , femur d 1-1-1, p d1ap; tibia v 2-2 -2; metatarsus v 2 bas. II , femur d 1-1-1, p and r d1ap; tibia v r1-r1-2; metatarsus v 2 bas. III , femur d 1-1-1, p 0-d1-d1 or d1ap, r d1ap; patella r d1; tibia v p1-p1-2, p 1-d1-1-0, r d1-1, d r1bas; metatarsus v 2 -0-comb, p and r d1-1-1, d 0-p1- 2 . IV , femur d 1-1-1, p and r d1ap; patella r d1; tibia v p1-2-2 or left 2-2-(p1-2), p 1-d1-1-0, r 1-d1-1-0 or left 1-d1-d1-1; metatarsus v 2-2 -comb or left r1-2-p1-comb, p and r d1-1-1 or left p d1-1-1-0-1, d 0-p1-2 (left leg seemingly abnormal, not considered for scoring of dataset). Opisthosoma length 2.17, spiracle–epigastrium 1.21, spiracle– spinnerets 0.24. Color in ethanol ( Fig. 8 ): As in female, but paler, specimen slightly faded. Palp ( Fig. 10 ): Tibia short, width/length 1.58, RTA longer than tibia, curved, compressed. Cymbial conductor wide. Tegulum displaced basally in ventral view, mostly visible in retrolateral view. Sperm duct with conspicuous loop on anterior-dorsal margin. Embolus long, with flat, rounded basal process. Median apophysis wide, truncate, translucent. Primary conductor with canal, ending in truncate sclerotized projection. Secondary conductor ventrally concave, well sclerotized, separated from anterior-dorsal sperm duct loop by a membranous stripe. Paramedian apophysis with prolateral-ventral cusp (pPMA, Fig. 23 d–f) and a series of several short cusps (rPMA). FIGURE 8. Coptoprepes bellavista new species . a–d male holotype ( a dorsal habitus; b prosoma, dorsal view; c ventral habitus, ventral view; d prosoma, ventral view). FIGURE 9. Coptoprepes bellavista new species . a–d female paratype ( a dorsal habitus; b prosoma, dorsal view; c ventral habitus, ventral view; d prosoma, ventral view). FIGURE 10. Coptoprepes bellavista new species . a–f male, g–h female paratype ( a left palp, male holotype, prolateral view, b same, ventral view, c same, retrolateral view; d copulatory bulb, preparation MJR-01329, prolateral view, e same, apical view, f same, retrolateral view; g epigyne, ventral view; h vulva). Female ( paratype ): Prosoma length 2.10, width 1.37. Length of tibia/metatarsus: I, 0.98/0.85; II, 0.88/ 0.82; III, 0.78/0.90; IV, 1.25/1.42. Chelicerae with five teeth on retromargin, four on promargin. Sternum length 1.05, width 0.83. Spines: leg I , femur d 1-1-1, p 2ap; tibia v 2-2 -p1; metatarsus v 2 bas. II , femur d 1-1- 1, p 1ap; tibia v r1-r1-p1; metatarsus v 2 bas. III , femur d 1-1-1, p and r 0-d1-d1; patella r d1; tibia v p1-p1-2, p 1-d1-1-0, r 1-d1-1-0 or 0-d1-1-0, d r1bas; metatarsus v 2 -0-comb, p and r d1-1-1, d 0-p1- 2 . IV , femur d 1-1- 1, p and r d1ap; patella r d1; tibia v p1-2-2, p and r 1-d1-1-0, d r1bas; metatarsus v 2-2 -2 plus apical comb, p d1-1-1, r d1-0-1, d 0-2-2. Color in ethanol ( Fig. 9 ): Prosoma brown, legs brown, darker distally. Opisthosoma pale brown with dorsal pattern brown, with weakly defined posterior chevrons. Opisthosoma length 2.70, epigastrium–spiracle 1.32, spiracle–spinnerets 0.18. Epigyne: Trapezoidal sclerotized patch, mostly anterior to internal ducts. Lateral lobes separate, posterior half not sclerotized except at the limits with the median field. Median field weakly sclerotized ( Fig. 10 g). Copulatory openings very close to epigastric furrow. Copulatory ducts not wrapped. Ducts of accessory bulbs short. Spermathecae with convoluted chambers, fertilization ducts well separated from posterior margin of epigyne ( Figs. 10 h, 24d–f). Variability: Spines in females: I, tibia v 2-2 -0. Spines in males: III, tibia v p1-2-2. Distribution: Known for Cautín and Ranco provinces in Chile . Other material examined: CHILE : Región IX (Araucanía): Cautín: same locality as holotype , 3 males , 2 females [plus 4 immatures] ( AMNH ); 15 km NE Villarrica, Flor del Lago, 300 m , 2 FITS , Nothofagus forest, 14 December 1984 – 10 February 1985 , S. & J. Peck, 1 male and 1 female ( AMNH ), 1male ( AMNH ; temporary preparation MJR-01329), 1 female ( AMNH ), 28 males , 7 females [plus 2 males without palps] ( AMNH ); Monte Verde, Carahue, 600 m , 30 January 1993 – 2 February 1993 , L. E. Peña, 1 male ( AMNH ; temporary preparation MJR-01348); 21 km NE Pucón, Lago Caburga, 600 m , FIT, mixed forest remnant, 15 December 1984 – 10 February 1985 , S. & J. Peck, 1 male and 3 females ( AMNH ); 10 km S Pucón, Volcán Villarrica Natl. Park, 900 m , FIT, Nothofagus grove on ash, 15 December 1984 – 10 February 1985 , S. & J. Peck, 3 males and 1 female ( AMNH ). Región XIV (Los Ríos): Ranco: 34 km WNW La Unión, 700m , FIT, mixed ever green forest, 17 December 1984 – 7 February 1985 , S. & J. Peck, 7 males and 3 females ( AMNH ).