A revision and phylogenetic analysis of the spider genus Coptoprepes Simon (Araneae: Anyphaenidae, Amaurobioidinae)
Author
Werenkraut, Victoria
Author
Ramírez, Martín J.
text
Zootaxa
2009
2212
1
40
journal article
10.5281/zenodo.275145
344a109d-a561-4054-97b7-e1850acc321f
1175-5326
275145
Coptoprepes recinto
new species
Figs. 19–21
,
22
f,
23m
–o,
24m
–o
Type
material:
Holotype
:
male and female
paratype
from
Chile
, Región VIII (Biobío), Ñuble province,
19.5 km
ESE
Recinto
, ca.
37º00’ S
,
71º30’ W
,
1250 m
, site 647, window trap,
Nothofagus
forest,
10 December 1982
–
3 January 1983
, A. Newton & M. Thayer (
AMNH
; temporary preparations, male: MJR-01335, VIW-00011, female: MJR-01339, VIW-00010).
Etymology:
The specific name is a noun in apposition taken from the
type
locality.
Diagnosis:
Females are similar to
C. nahuelbuta
by having the epigynal median field hidden in the epigastric furrow, but can be distinguished by the notched posterior epigynal margin (
Figs.
21
g–h,
24m
–n). Males are distinguished by the long and curved RTA, about twice as long as the palpal tibia, and the flat, rounded and projecting median apophysis (
Figs. 21
c–f,
23m
–o).
Description:
Male
(
holotype
): Prosoma length 1.92, width 1.34, anteriorly slightly narrower than in female. Length of tibia/metatarsus: I, 1.24/1.14; II, 1.05/1.02; III, 0.81/0.99; IV, 1.18/1.45. Chelicerae less robust than in female, with five teeth on retromargin, four on promargin. Sternum length 1.08, width 0.84. Spines: leg
I
, femur d 1-1-1, p 2ap; tibia
v 2-2
-p1; metatarsus
v 2
bas.
II
, femur d 1-1-1, p d1ap; tibia v r1-r1- p1, p 0-1 or 0; metatarsus
v 2
bas.
III
, femur d 1-1-1, p 0-d1-d1, r d1ap; patella r d1; tibia v p1-p1-2, d r1bas, p 1-d1-1-0, r d1-1; metatarsus
v 2
bas plus apical comb, d 0-2-2, p and r d1-1-
1.
IV
, femur d 1-1-1, p and r d1ap; patella r d1; tibia v p1-2-p1 or p1-2-2, d r1bas, p and r 1-d1-1-0; metatarsus
v 2-2
-0 plus apical comb, d 0-2-2, p and r d1-1-1. Opisthosoma length 2.22, spiracle–epigastrium 1.29, spiracle–spinnerets 0.11. Color in ethanol (
Fig. 19
): Prosoma and legs dark brown, almost uniform, except coxae pale brown, and a pale apical patch ventrally on leg femora III and IV. Opisthosoma dark brown with paler pattern, dorsum with anterior lateral bands of white setae and posterior chevrons. Palp (
Fig. 21
): Tibia width/length 1.42, RTA long, about twice as long as tibia, curved, compressed. Cymbial conductor wide, conspicuous. Tegulum basal/retrolateral in ventral view. Sperm duct with two conspicuous loops on anterior-dorsal margin, small apical tegular sclerite with conical shape arising between both loops. Embolus long, thin, basal process ample, flattened, slightly projecting with rounded border. Median apophysis flat, concave, projecting anteriorly, with prolateral-dorsal acute projection. Primary conductor with canal, ending in acute sclerotized projection. Secondary conductor a small sclerotized cone, separated from anterior-dorsal sperm duct loop by membranous stripe. Paramedian apophysis reduced to sclerotized stripe connecting with base of median apophysis, with two rounded flat cusps (
Fig.
23
m–o).
FIGURE 19.
Coptoprepes recinto
new species
.
a–d
male holotype (
a
dorsal habitus;
b
prosoma, dorsal view;
c
ventral habitus, ventral view;
d
prosoma, ventral view).
FIGURE 20.
Coptoprepes recinto
new species
.
a–d
female paratype (
a
dorsal habitus;
b
prosoma, dorsal view;
c
ventral habitus, ventral view;
d
prosoma, ventral view).
FIGURE 21.
Coptoprepes recinto
new species
.
a–f
male,
g–h
female paratype (
a
left palp, male holotype, prolateral view,
b
same, ventral view,
c
same, retrolateral view;
d
copulatory bulb, preparation MJR-01332, prolateral view,
e
same, apical view,
f
same, retrolateral view;
g
epigyne, ventral view;
h
vulva).
Female
(
paratype
): Prosoma length 2.05, width 1.37, wider between legs II–III. Tibia/metatarsus length: I, 1.65/0.89; II, 0.93/0.89; III, 0.8/0.97; IV, 1.22/1.47. Chelicerae with four teeth on promargin, six on retromargin. Sternum length 1.15, width 0.89. Spines: leg
I
, femur d 1-1-1, p 2ap; tibia
v 2-2
-0 or r1-2-0; metatarsus
v 2
bas.
II
, femur d 1-1-1, p d1ap; tibia v r1-r1-p1; metatarsus
v 2
bas.
III
, femur d 1-1-1, p 0-d1-d1, r d1 ap; patella r d1; tibia v 0-p1-2 or p1-p1-2, d r1bas, p 1-d1-1-0, r d1-1; metatarsus
v 2
-0-0 plus apical comb, d 0-2-2, p and r d1-1-
1.
IV
, femur d 1-1-1, p d1ap, r d1ap; patella r d1; tibia v p1-2-2, d r1bas, p and r 1- d1-1-0; metatarsus
v 2-2
-0 plus apical comb, d 0-2-2, p and r d1-1-1. Color in ethanol (
Fig. 20
): As in male, but in poor condition. Opisthosoma length 3.40, epigastrium–spiracle approximately 1.60, spiracle–spinnerets 0.33, in poor condition. Epigyne (
Fig.
21
g): Ventrally a single plate with a posterior notch, lateral lobes well separated in posterior-dorsal view, their limits with median field parallel. Copulatory ducts running in ectal arcs. Ducts of accessory bulbs short. Spermathecae with convoluted chambers (
Fig. 21
h).
Variability:
Spines in males: III, tibia v p1-2-2. Spines in females: I, tibia
v 2-2
-p1.
Distribution:
Known for Ñuble and Malleco provinces in
Chile
.
Other material examined:
CHILE
:
Región VIII (Biobío):
Ñuble: same locality as
holotype
,
12 males
,
4 females
[plus 2 immature males and 2 immatures];
22.7 km
ESE
Recinto,
1330 m
, site 646, window trap,
Nothofagus
forest,
10 December 1982
–
3 January 1983
, A. Newton & M. Thayer,
1 male
(
AMNH
; temporary preparation MJR-01332),
3 males
(
AMNH
);
72 km
SE Chillán, Trancas, nr. Termas,
1700 m
, FIT,
Nothofagus
forest,
6 December 1984
–
19 February 1985
, S. & J. Peck,
3 males
and
1 female
(
AMNH
).
Región IX (Araucanía):
Malleco province:
12 km
E Malalcahuello,
1350 m
, site 650, window trap,
Nothofagus dombeyi
/
Araucaria
forest,
13–31 December1982
, A. Newton & M. Thayer,
3 males
(
AMNH
);
14 km
E Malalcahuello,
1570 m
, site 649, window trap,
Nothofagus
pumilio-Araucaria
forest,
13–31 December 1982
, A. Newton & M. Thayer,
1 female
(
AMNH
).