Order Chiroptera - Family Pteropodidae Author Wilson, Don E. Author Reeder, DeeAnn text 2005 The Johns Hopkins University Press Baltimore Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1 313 350 book chapter 0-8018-8221-4 10.5281/zenodo.7316519 Paranyctimene tenax subsp. tenax Bergmans 2001 Paranyctimene tenax subsp. tenax Bergmans 2001 , Beaufortia, 51: 146 . Type Locality: Papua New Guinea , Morobe Province , 32 km SSW of Wau, upstream of Anadea, about 07°36'S , 146°37'E , 850 m .