<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFFBFFBB5AB6F99B4352CD49" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus naikii V. S. Dang &amp; Vuong 2022" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
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<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571816" ID-Zenodo-Dep="7571816" LSID="urn:lsid:plazi:treatment:43284262FFFBFFBB5AB6F99B4352CD49" httpUri="http://treatment.plazi.org/id/43284262FFFBFFBB5AB6F99B4352CD49" lastPageId="101" pageId="100">
<subSubSection box="[136,1084,1598,1625]" pageId="100" type="nomenclature">
<paragraph blockId="100.[136,1452,1598,1732]" box="[136,1084,1598,1625]" pageId="100">
<emphasis bold="true" box="[136,171,1598,1624]" pageId="100">49.</emphasis>
<taxonomicName authority="V. S. Dang &amp; Vuong (2022: 20)" authorityName="V. S. Dang &amp; Vuong" authorityPageNumber="20" authorityYear="2022" box="[178,729,1598,1625]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="100" phylum="Tracheophyta" rank="species" species="naikii">
<emphasis bold="true" box="[178,381,1598,1625]" italics="true" pageId="100">Lasianthus naikii</emphasis>
V.S.Dang &amp; Vuong (2022: 20)
</taxonomicName>
— 
<figureCitation box="[759,837,1598,1624]" captionStart="FIGURE 76" captionStartId="102.[136,229,1778,1800]" captionTargetBox="[167,1418,194,1753]" captionTargetId="figure-26@102.[165,1418,193,1754]" captionTargetPageId="102" captionText="FIGURE 76. Lasianthus naikii V.S.Dang &amp; Vuong (from Dang 453, VNM): A. Branche, B. Stipules, C. Abaxial leaf surfaces, D. Calyces, E–F. Close-up of flowers, G–H. Mature fruits, I. Seeds." figureDoi="http://doi.org/10.5281/zenodo.7571786" httpUri="https://zenodo.org/record/7571786/files/figure.png" pageId="100">Fig. 76</figureCitation>
(plate), 
<figureCitation box="[931,1010,1598,1624]" captionStart="FIGURE 73" captionStartId="98.[136,229,1801,1823]" captionTargetBox="[173,1400,200,1757]" captionTargetId="figure-24@98.[165,1418,193,1778]" captionTargetPageId="98" captionText="FIGURE 73. Distribution maps of Lasianthus longissimus, L. lucidus, L. micranthus and L. naikii." figureDoi="http://doi.org/10.5281/zenodo.7571776" httpUri="https://zenodo.org/record/7571776/files/figure.png" pageId="100">Fig. 73</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection pageId="100" type="materials_examined">
<paragraph blockId="100.[136,1452,1598,1732]" pageId="100">
<emphasis bold="true" box="[136,194,1638,1659]" pageId="100">Type:</emphasis>
— 
<materialsCitation collectingDate="2021-04-22" collectionCode="VNM, HN" collectorName="Van-Son Dang &amp; Ba Vuong Truong &amp; Minh Quan Dang &amp; Nghia Son Hoang &amp; Dang" country="Vietnam" county="K'Bang District" elevation="1050" location="Kon Chu Rang Nature Reserve" municipality="Son Lang Commune" pageId="100" specimenCode="VNM00042916, VNM00042917, VNM00042918" specimenCount="1" stateProvince="Gia Lai" typeStatus="holotype">
<collectingCountry box="[219,332,1638,1659]" name="Vietnam" pageId="100">VIETNAM</collectingCountry>
. 
<collectingRegion box="[343,513,1638,1660]" country="Vietnam" name="Gia Lai" pageId="100">Gia Lai Province</collectingRegion>
, 
<collectingCounty box="[524,682,1638,1660]" pageId="100">K’Bang District</collectingCounty>
, 
<collectingMunicipality box="[693,898,1638,1659]" pageId="100">Son Lang Commune</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:43284262FFFBFFBB5AB6F99B4352CD49:CE5EA5AFFFFBFFBA59B3F9C344E7CA33" box="[909,1217,1638,1660]" country="Vietnam" county="K'Bang District" municipality="Son Lang Commune" name="Kon Chu Rang Nature Reserve" pageId="100" stateProvince="Gia Lai">Kon Chu Rang Nature Reserve</location>
, 
<quantity box="[1228,1362,1638,1659]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="100" unit="m" value="1050.0" valueMax="1100.0" valueMin="1000.0">
<elevation box="[1228,1362,1638,1659]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="100" unit="m" value="1050.0" valueMax="1100.0" valueMin="1000.0">1000–1100 m</elevation>
</quantity>
elev., 
<date pageId="100" value="2021-04-22">
<collectingDate pageId="100" value="2021-04-22">22 April 2021</collectingDate>
</date>
, 
<emphasis box="[304,1106,1674,1695]" italics="true" pageId="100">
<collectorName box="[304,446,1674,1695]" pageId="100">Van-Son Dang</collectorName>
, 
<collectorName box="[454,624,1674,1695]" pageId="100">Ba Vuong Truong</collectorName>
, 
<collectorName box="[633,800,1674,1695]" pageId="100">Minh Quan Dang</collectorName>
&amp; 
<collectorName box="[829,1002,1674,1695]" pageId="100">Nghia Son Hoang</collectorName>
, 
<collectorName box="[1011,1064,1674,1695]" pageId="100">Dang</collectorName>
453
</emphasis>
(
<typeStatus box="[1117,1203,1674,1696]" pageId="100">holotype</typeStatus>
<collectionCode box="[1207,1268,1674,1695]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="100" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[1282,1443,1674,1696]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="100" type="Herbarium">VNM00042916</specimenCode>
]; isotypes 
<collectionCode box="[274,315,1710,1731]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="100" type="Herbarium">HN</collectionCode>
!, 
<collectionCode box="[328,389,1710,1731]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="100" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[404,560,1710,1731]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="100" type="Herbarium">VNM00042917</specimenCode>
, 
<specimenCode box="[570,729,1710,1731]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="100" type="Herbarium">VNM00042918</specimenCode>
])
</materialsCitation>
.
</paragraph>
<paragraph blockId="100.[136,1452,1778,2095]" pageId="100">
<emphasis bold="true" box="[136,396,1778,1804]" pageId="100">
<collectingRegion box="[136,262,1778,1804]" pageId="100">Specimens</collectingRegion>
examined:
</emphasis>
— 
<materialsCitation collectingDate="2021-04-25" collectionCode="VNM" collectorName="Van-Son Dang &amp; Ba Vuong Truong &amp; Minh Quan Dang &amp; Nghia Son Hoang &amp; Dang" country="Vietnam" county="K'Bang District" elevation="1050" location="Kon Chu Rang Nature Reserve" municipality="Son Lang Commune" pageId="100" specimenCount="1" stateProvince="Gia Lai">
<collectingCountry box="[425,557,1778,1804]" name="Vietnam" pageId="100">VIETNAM</collectingCountry>
. 
<emphasis bold="true" box="[570,671,1778,1804]" pageId="100">Central:</emphasis>
<collectingRegion box="[681,880,1778,1804]" country="Vietnam" name="Gia Lai" pageId="100">Gia Lai Province</collectingRegion>
, 
<collectingCounty box="[894,1079,1778,1804]" pageId="100">K’Bang District</collectingCounty>
, 
<collectingMunicipality box="[1093,1333,1778,1805]" pageId="100">Son Lang Commune</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:43284262FFFBFFBB5AB6F99B4352CD49:CE5EA5AFFFFBFFBA5F7DF956415CCB79" country="Vietnam" county="K'Bang District" municipality="Son Lang Commune" name="Kon Chu Rang Nature Reserve" pageId="100" stateProvince="Gia Lai">Kon Chu Rang Nature Reserve</location>
, 
<quantity box="[389,474,1814,1841]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="100" unit="m" value="1050.0">
<elevation box="[389,474,1814,1841]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="100" unit="m" value="1050.0">1050 m</elevation>
</quantity>
elev., 
<date box="[546,704,1814,1841]" pageId="100" value="2021-04-25">
<collectingDate box="[546,704,1814,1841]" pageId="100" value="2021-04-25">25 April 2021</collectingDate>
</date>
, 
<emphasis italics="true" pageId="100">
<collectorName box="[715,880,1815,1841]" pageId="100">Van-Son Dang</collectorName>
, 
<collectorName box="[891,1091,1815,1840]" pageId="100">Ba Vuong Truong</collectorName>
, 
<collectorName box="[1102,1298,1814,1841]" pageId="100">Minh Quan Dang</collectorName>
&amp; 
<collectorName pageId="100">Nghia Son Hoang</collectorName>
, 
<collectorName box="[226,288,1851,1876]" pageId="100">Dang</collectorName>
453a
</emphasis>
(
<collectionCode box="[367,438,1850,1876]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="100" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="101" pageId="100" type="distribution">
<paragraph blockId="100.[136,1452,1778,2095]" pageId="100">
<emphasis bold="true" box="[189,345,1886,1912]" pageId="100">Distribution:</emphasis>
—Endemic to 
<collectingCountry box="[520,615,1886,1912]" name="Vietnam" pageId="100">Vietnam</collectingCountry>
(currently known only at Kon Chu Rang Nature Reserve of 
<collectingRegion country="Vietnam" name="Gia Lai" pageId="100">Gia Lai Province</collectingRegion>
).
</paragraph>
<paragraph blockId="100.[136,1452,1778,2095]" pageId="100">
<emphasis bold="true" box="[189,422,1958,1984]" pageId="100">Habitat &amp; Ecology:</emphasis>
— 
<taxonomicName authorityName="V. S. Dang &amp; Vuong" authorityYear="2022" box="[450,643,1958,1984]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="100" phylum="Tracheophyta" rank="species" species="naikii">
<emphasis box="[450,643,1958,1984]" italics="true" pageId="100">Lasianthus naikii</emphasis>
</taxonomicName>
grows under the shade of the primary evergreen forest at an elevation of 
<quantity box="[136,293,1994,2021]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="100" unit="m" value="1050.0" valueMax="1100.0" valueMin="1000.0">1000–1100 m</quantity>
.
</paragraph>
<paragraph blockId="100.[136,1452,1778,2095]" box="[189,897,2030,2057]" pageId="100">
<emphasis bold="true" box="[189,323,2030,2056]" pageId="100">Phenology:</emphasis>
—Flowering and fruiting from February to August.
</paragraph>
<paragraph blockId="100.[136,1452,1778,2095]" box="[189,617,2066,2095]" pageId="100">
<emphasis bold="true" box="[189,405,2066,2092]" pageId="100">Vernacular name:</emphasis>
—Xú hưƠng naiki.
</paragraph>
<paragraph blockId="101.[136,1451,158,257]" pageId="101">
<emphasis bold="true" box="[189,265,158,184]" pageId="101">Notes:</emphasis>
— 
<taxonomicName authorityName="V. S. Dang &amp; Vuong" authorityYear="2022" box="[294,486,158,184]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="101" phylum="Tracheophyta" rank="species" species="naikii">
<emphasis box="[294,486,158,184]" italics="true" pageId="101">Lasianthus naikii</emphasis>
</taxonomicName>
was described from a Vietnamese specimen (
<emphasis box="[989,1171,158,184]" italics="true" pageId="101">Dang 453, VNM</emphasis>
) by 
<bibRefCitation author="Dang, M. Q. &amp; Nguyen, M. H. &amp; Hoang, N. S. &amp; Le, V. T. &amp; Nguyen, T. M. H. &amp; Ho, N. Q. C. &amp; Truong, B. V. &amp; Dang, V. S." box="[1220,1425,158,185]" pageId="101" pagination="19 - 31" refId="ref53709" refString="Dang, M. Q., Nguyen, M. H., Hoang, N. S., Le, V. T., Nguyen, T. M. H., Ho, N. Q. C., Truong, B. V. &amp; Dang, V. S. (2022) Two new species of Lasianthus Jack (Rubiaceae) from southern Vietnam. European Journal of Taxonomy 806: 19 - 31. https: // doi. org / 10.5852 / ejt. 2022.806.1697" type="journal article" year="2022">
Dang 
<emphasis box="[1286,1341,158,184]" italics="true" pageId="101">et al.</emphasis>
(2022)
</bibRefCitation>
. It is similar to 
<taxonomicName authority="Matsum. (1901: 17)" authorityName="Matsum." authorityPageNumber="17" authorityYear="1901" box="[276,672,194,221]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="101" phylum="Tracheophyta" rank="species" species="formosensis">
<emphasis box="[276,439,194,220]" italics="true" pageId="101">L. formosensis</emphasis>
Matsum. (1901: 17)
</taxonomicName>
, but differs from the latter in having less than secondary veins, bracts present (vs. absent), longer calyx tubes, and corolla lobes 6 (vs. 5).
</paragraph>
</subSubSection>
</treatment>
</document>