<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFE8FFA95AB6FD7B4547C95C" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus saprosmoides Pit. 1924" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571870" ID-Zenodo-Dep="7571870" LSID="urn:lsid:plazi:treatment:43284262FFE8FFA95AB6FD7B4547C95C" httpUri="http://treatment.plazi.org/id/43284262FFE8FFA95AB6FD7B4547C95C" pageId="119">
<subSubSection box="[136,1008,734,761]" pageId="119" type="nomenclature">
<paragraph blockId="119.[136,1452,734,832]" box="[136,1008,734,761]" pageId="119">
<emphasis bold="true" box="[136,171,734,760]" pageId="119">60.</emphasis>
<taxonomicName ID-CoL="3SCS6" authority="Pit. (1924: 387)" authorityName="Pit." authorityPageNumber="387" authorityYear="1924" box="[178,653,734,761]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="119" phylum="Tracheophyta" rank="species" species="saprosmoides">
<emphasis bold="true" box="[178,470,734,761]" italics="true" pageId="119">Lasianthus saprosmoides</emphasis>
Pit. (1924: 387)
</taxonomicName>
— 
<figureCitation box="[683,761,734,760]" captionStart="FIGURE 90" captionStartId="120.[136,229,1789,1811]" captionTargetBox="[169,1415,194,1756]" captionTargetId="figure-24@120.[166,1418,193,1765]" captionTargetPageId="120" captionText="FIGURE 90. Lasianthus saprosmoides Pit. (from Dang 429, VNM): A. Branch with fruits, B. Stipules, C. Abaxial leaf surfaces, D–F. Inflorescence showing side view buds and flower, G–I. Young fruits." figureDoi="http://doi.org/10.5281/zenodo.7571830" httpUri="https://zenodo.org/record/7571830/files/figure.png" pageId="119">Fig. 90</figureCitation>
(plate), 
<figureCitation box="[855,934,734,760]" captionStart="FIGURE 89" captionStartId="118.[136,229,1814,1836]" captionTargetBox="[176,1405,203,1775]" captionTargetId="figure-24@118.[165,1418,193,1790]" captionTargetPageId="118" captionText="FIGURE 89. Distribution maps of Lasianthus sapaensis, L. saprosmoides, L. saxorum and L. schmidtii." figureDoi="http://doi.org/10.5281/zenodo.7571826" httpUri="https://zenodo.org/record/7571826/files/figure.png" pageId="119">Fig. 89</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection pageId="119" type="materials_examined">
<paragraph blockId="119.[136,1452,734,832]" pageId="119">
<emphasis bold="true" box="[136,194,774,795]" pageId="119">Type:</emphasis>
— 
<materialsCitation collectingDate="1919-04-28" collectorName="A. Chevalier" country="Vietnam" elevation="1500" location="Langbian Plateau" pageId="119" specimenCount="1" stateProvince="Lam Dong" typeStatus="holotype">
<collectingCountry box="[219,332,774,795]" name="Vietnam" pageId="119">VIETNAM</collectingCountry>
. 
<collectingRegion box="[349,563,774,796]" country="Vietnam" name="Lam Dong" pageId="119">Lam Dong Province</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:43284262FFE8FFA95AB6FD7B4547C95C:CE5EA5AFFFE8FFA9587AFCA342D0CF54" box="[580,758,774,796]" country="Vietnam" name="Langbian Plateau" pageId="119" stateProvince="Lam Dong">Langbian Plateau</location>
, 
<quantity box="[776,857,774,796]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="119" unit="m" value="1500.0">
<elevation box="[776,857,774,796]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="119" unit="m" value="1500.0">1500 m</elevation>
</quantity>
elev., 
<date box="[936,1088,774,796]" pageId="119" value="1919-04-28">
<collectingDate box="[936,1088,774,796]" pageId="119" value="1919-04-28">28 April 1919</collectingDate>
</date>
, 
<emphasis box="[1106,1310,774,795]" italics="true" pageId="119">
<collectorName box="[1106,1236,774,795]" pageId="119">A. Chevalier</collectorName>
40329
</emphasis>
(
<typeStatus box="[1330,1416,774,796]" pageId="119">holotype</typeStatus>
P! [P02285236])
</materialsCitation>
.
</paragraph>
<paragraph blockId="119.[136,1452,878,1301]" pageId="119">
<emphasis bold="true" box="[136,394,878,904]" pageId="119">
<collectingRegion box="[136,262,878,904]" pageId="119">Specimens</collectingRegion>
examined:
</emphasis>
— 
<materialsCitation collectingDate="1919-04-28" collectorName="A. Chevalier" country="Vietnam" elevation="1500" location="Langbian Plateau" pageId="119" specimenCount="1" stateProvince="Lam Dong">
<collectingCountry box="[423,555,878,904]" name="Vietnam" pageId="119">VIETNAM</collectingCountry>
. 
<emphasis bold="true" box="[566,667,878,904]" pageId="119">Central:</emphasis>
<collectingRegion box="[675,905,878,905]" country="Vietnam" name="Lam Dong" pageId="119">Lam Dong Province</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:43284262FFE8FFA95AB6FD7B4547C95C:CE5EA5AFFFE8FFA959A8FCCB447ACFC0" box="[918,1116,878,904]" country="Vietnam" name="Langbian Plateau" pageId="119" stateProvince="Lam Dong">Langbian Plateau</location>
, 
<quantity box="[1128,1214,878,905]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="119" unit="m" value="1500.0">
<elevation box="[1128,1214,878,905]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="119" unit="m" value="1500.0">1500 m</elevation>
</quantity>
elev., 
<date box="[1287,1447,878,904]" pageId="119" value="1919-04-28">
<collectingDate box="[1287,1447,878,904]" pageId="119" value="1919-04-28">28 April 1919</collectingDate>
</date>
, 
<emphasis box="[136,353,914,941]" italics="true" pageId="119">
<collectorName box="[136,276,914,941]" pageId="119">A. Chevalier</collectorName>
40329
</emphasis>
(P); in the
</materialsCitation>
<materialsCitation collectingDate="2019-10-20" collectionCode="VNM" collectorName="Dang V. S. &amp; Dang" country="Vietnam" elevation="1405" location="Bidoup Nui Ba National Park" pageId="119" specimenCount="1" stateProvince="Central">
same location, 
<location LSID="urn:lsid:plazi:treatment:43284262FFE8FFA95AB6FD7B4547C95C:CE5EA5AFFFE8FFA958BFFC3743EDCFE4" box="[641,971,914,941]" country="Vietnam" name="Bidoup Nui Ba National Park" pageId="119" stateProvince="Central">Bidoup Nui Ba National Park</location>
, 
<quantity box="[981,1065,914,941]" metricMagnitude="3" metricUnit="m" metricValue="1.405" pageId="119" unit="m" value="1405.0">
<elevation box="[981,1065,914,941]" metricMagnitude="3" metricUnit="m" metricValue="1.405" pageId="119" unit="m" value="1405.0">1405 m</elevation>
</quantity>
elev., 
<date box="[1136,1325,914,940]" pageId="119" value="2019-10-20">
<collectingDate box="[1136,1325,914,940]" pageId="119" value="2019-10-20">20 October 2019</collectingDate>
</date>
, 
<emphasis italics="true" pageId="119">
<collectorName box="[1335,1447,915,941]" pageId="119">Dang V.S.</collectorName>
, 
<collectorName box="[136,198,951,976]" pageId="119">Dang</collectorName>
429
</emphasis>
(
<collectionCode box="[262,333,950,976]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="119" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="119" type="distribution">
<paragraph blockId="119.[136,1452,878,1301]" pageId="119">
<emphasis bold="true" box="[189,345,986,1012]" pageId="119">Distribution:</emphasis>
—Endemic to 
<collectingCountry box="[513,608,986,1012]" name="Vietnam" pageId="119">Vietnam</collectingCountry>
(currently known only from Bidoup Nui Ba National Park of 
<collectingRegion country="Vietnam" name="Lam Dong" pageId="119">Lam Dong Province</collectingRegion>
).
</paragraph>
<paragraph blockId="119.[136,1452,878,1301]" pageId="119">
<emphasis bold="true" box="[189,426,1058,1084]" pageId="119">Habitat &amp; Ecology:</emphasis>
—Growing in the understory of primary and secondary forests or the hill of the evergreen montane forest at 
<quantity box="[337,492,1094,1121]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.7" metricValueMin="1.0" pageId="119" unit="m" value="1350.0" valueMax="1700.0" valueMin="1000.0">1000–1700 m</quantity>
elev.
</paragraph>
<paragraph blockId="119.[136,1452,878,1301]" box="[189,891,1130,1157]" pageId="119">
<emphasis bold="true" box="[189,323,1130,1156]" pageId="119">Phenology:</emphasis>
—Flowering and fruiting from April to November.
</paragraph>
<paragraph blockId="119.[136,1452,878,1301]" box="[189,609,1166,1195]" pageId="119">
<emphasis bold="true" box="[189,405,1166,1192]" pageId="119">Vernacular name:</emphasis>
—Xú hưƠng nhại.
</paragraph>
<paragraph blockId="119.[136,1452,878,1301]" pageId="119">
<emphasis bold="true" box="[189,265,1202,1228]" pageId="119">Notes:</emphasis>
— 
<taxonomicName box="[294,578,1202,1228]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="119" phylum="Tracheophyta" rank="species" species="saprosmoides">
<emphasis box="[294,578,1202,1228]" italics="true" pageId="119">Lasianthus saprosmoides</emphasis>
</taxonomicName>
was described as new from a Vietnamese specimen (
<emphasis box="[1185,1406,1202,1229]" italics="true" pageId="119">A. Chevalier 40329</emphasis>
) by 
<bibRefCitation author="Pitard, J." box="[136,290,1238,1264]" pageId="119" pagination="395" refId="ref55116" refString="Pitard, J. (1924) Rubiacees In: Lecomte, P. H., Humbert, H. &amp; Gagnepain, F. (Eds.) Flore generale de l'Indo-Chine 3: 395. Masson, Paris, pp. 20 - 442." type="journal article" year="1924">Pitard (1924)</bibRefCitation>
. It is characterized in having oblong or ovate-lanceolate leaves, secondary veins 5–6 on each side of midrib, calyx tube truncate and calyx lobes barely visible, corolla lobes 3, and fruits globose, 
<quantity box="[1185,1278,1274,1301]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="119" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
in diam.
</paragraph>
</subSubSection>
</treatment>
</document>