<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFE4FFA25AB6F99B4080CD6D" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus sonlangensis V. S. Dang, Vuong &amp; Quan 2022" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571888" ID-Zenodo-Dep="7571888" LSID="urn:lsid:plazi:treatment:43284262FFE4FFA25AB6F99B4080CD6D" httpUri="http://treatment.plazi.org/id/43284262FFE4FFA25AB6F99B4080CD6D" lastPageId="124" pageId="123">
<subSubSection box="[136,1241,1598,1625]" pageId="123" type="nomenclature">
<paragraph blockId="123.[136,1452,1598,1732]" box="[136,1241,1598,1625]" pageId="123">
<emphasis bold="true" box="[136,171,1598,1624]" pageId="123">64.</emphasis>
<taxonomicName authority="V. S. Dang, Vuong &amp; Quan (2022: 25)" authorityName="V. S. Dang, Vuong &amp; Quan" authorityPageNumber="25" authorityYear="2022" box="[178,885,1598,1625]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="123" phylum="Tracheophyta" rank="species" species="sonlangensis">
<emphasis bold="true" box="[178,463,1598,1625]" italics="true" pageId="123">Lasianthus sonlangensis</emphasis>
V.S.Dang, Vuong &amp; Quan (2022: 25)
</taxonomicName>
— 
<figureCitation box="[916,994,1598,1625]" captionStart="FIGURE 95" captionStartId="126.[136,229,1784,1806]" captionTargetBox="[167,1418,194,1758]" captionTargetId="figure-24@126.[165,1418,193,1760]" captionTargetPageId="126" captionText="FIGURE 95. Lasianthus sonlangensis V.S.Dang, Vuong &amp; Quan (from Dang 465, VNM): A. Branche, B. Stipules, C. Adaxial and abaxial leaf surfaces, D–F. Close-up of infructescence, G–I. Close-up of flowers (photos by Truong Ba Vuong)." figureDoi="http://doi.org/10.5281/zenodo.7571844" httpUri="https://zenodo.org/record/7571844/files/figure.png" pageId="123">Fig. 95</figureCitation>
(plate), 
<figureCitation box="[1088,1167,1598,1624]" captionStart="FIGURE 94" captionStartId="125.[136,229,1814,1836]" captionTargetBox="[176,1405,205,1773]" captionTargetId="figure-17@125.[165,1418,193,1790]" captionTargetPageId="125" captionText="FIGURE 94. Distribution maps of Lasianthus sikkimensis, L. sonlangensis, L. stephanocalycinus and L. tamdaoensis." figureDoi="http://doi.org/10.5281/zenodo.7571842" httpUri="https://zenodo.org/record/7571842/files/figure.png" pageId="123">Fig. 94</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection pageId="123" type="materials_examined">
<paragraph blockId="123.[136,1452,1598,1732]" pageId="123">
<emphasis bold="true" box="[136,194,1638,1659]" pageId="123">Type:</emphasis>
— 
<materialsCitation collectingDate="2021-04-22" collectionCode="VNM, HN" collectorName="Van-Son Dang &amp; Ba Vuong Truong &amp; Minh Quan Dang &amp; Nghia Son Hoang &amp; Dang" country="Vietnam" county="K'Bang District" elevation="1000" location="Kon Chu Rang Nature Reserve" municipality="Son Lang Commune" pageId="123" specimenCode="VNM00042935, VNM00042936, VNM00042937" specimenCount="1" stateProvince="Gia Lai" typeStatus="holotype">
<collectingCountry box="[219,332,1638,1659]" name="Vietnam" pageId="123">VIETNAM</collectingCountry>
. 
<collectingRegion box="[343,515,1638,1660]" country="Vietnam" name="Gia Lai" pageId="123">Gia Lai Province</collectingRegion>
, 
<collectingCounty box="[527,686,1638,1660]" pageId="123">K’Bang District</collectingCounty>
, 
<collectingMunicipality box="[697,904,1638,1659]" pageId="123">Son Lang Commune</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:43284262FFE4FFA25AB6F99B4080CD6D:CE5EA5AFFFE4FFA559ADF9C344ECCA33" box="[915,1226,1638,1660]" country="Vietnam" county="K'Bang District" municipality="Son Lang Commune" name="Kon Chu Rang Nature Reserve" pageId="123" stateProvince="Gia Lai">Kon Chu Rang Nature Reserve</location>
, 
<quantity box="[1238,1360,1638,1659]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="123" unit="m" value="1000.0" valueMax="1100.0" valueMin="900.0">
<elevation box="[1238,1360,1638,1659]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="123" unit="m" value="1000.0" valueMax="1100.0" valueMin="900.0">900–1100 m</elevation>
</quantity>
elev., 
<date pageId="123" value="2021-04-22">
<collectingDate pageId="123" value="2021-04-22">22 April 2021</collectingDate>
</date>
, 
<emphasis box="[304,1106,1674,1695]" italics="true" pageId="123">
<collectorName box="[304,446,1674,1695]" pageId="123">Van-Son Dang</collectorName>
, 
<collectorName box="[454,624,1674,1695]" pageId="123">Ba Vuong Truong</collectorName>
, 
<collectorName box="[633,800,1674,1695]" pageId="123">Minh Quan Dang</collectorName>
&amp; 
<collectorName box="[829,1002,1674,1695]" pageId="123">Nghia Son Hoang</collectorName>
, 
<collectorName box="[1011,1064,1674,1695]" pageId="123">Dang</collectorName>
465
</emphasis>
(
<typeStatus box="[1117,1203,1674,1696]" pageId="123">holotype</typeStatus>
<collectionCode box="[1207,1268,1674,1695]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="123" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[1282,1442,1674,1696]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="123" type="Herbarium">VNM00042935</specimenCode>
]; isotypes 
<collectionCode box="[274,315,1710,1731]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="123" type="Herbarium">HN</collectionCode>
!, 
<collectionCode box="[328,389,1710,1731]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="123" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[404,561,1710,1732]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="123" type="Herbarium">VNM00042936</specimenCode>
, 
<specimenCode box="[570,729,1710,1731]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="123" type="Herbarium">VNM00042937</specimenCode>
])
</materialsCitation>
.
</paragraph>
<paragraph blockId="123.[136,1452,1778,2095]" pageId="123">
<emphasis bold="true" box="[136,396,1778,1804]" pageId="123">
<collectingRegion box="[136,262,1778,1804]" pageId="123">Specimens</collectingRegion>
examined:
</emphasis>
— 
<materialsCitation collectingDate="2021-04-25" collectionCode="VNM" collectorName="Van-Son Dang &amp; Ba Vuong Truong &amp; Minh Quan Dang &amp; Nghia Son Hoang &amp; Dang" country="Vietnam" county="K'Bang District" elevation="1080" location="Kon Chu Rang Nature Reserve" municipality="Son Lang Commune" pageId="123" specimenCount="1" stateProvince="Gia Lai">
<collectingCountry box="[425,557,1778,1804]" name="Vietnam" pageId="123">VIETNAM</collectingCountry>
. 
<emphasis bold="true" box="[570,671,1778,1804]" pageId="123">Central:</emphasis>
<collectingRegion box="[681,880,1778,1804]" country="Vietnam" name="Gia Lai" pageId="123">Gia Lai Province</collectingRegion>
, 
<collectingCounty box="[894,1079,1778,1804]" pageId="123">K’Bang District</collectingCounty>
, 
<collectingMunicipality box="[1093,1333,1778,1805]" pageId="123">Son Lang Commune</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:43284262FFE4FFA25AB6F99B4080CD6D:CE5EA5AFFFE4FFA55F7DF956415CCB79" country="Vietnam" county="K'Bang District" municipality="Son Lang Commune" name="Kon Chu Rang Nature Reserve" pageId="123" stateProvince="Gia Lai">Kon Chu Rang Nature Reserve</location>
, 
<quantity box="[389,474,1814,1841]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="123" unit="m" value="1080.0">
<elevation box="[389,474,1814,1841]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="123" unit="m" value="1080.0">1080 m</elevation>
</quantity>
elev., 
<date box="[546,704,1814,1841]" pageId="123" value="2021-04-25">
<collectingDate box="[546,704,1814,1841]" pageId="123" value="2021-04-25">25 April 2021</collectingDate>
</date>
, 
<emphasis italics="true" pageId="123">
<collectorName box="[715,880,1815,1841]" pageId="123">Van-Son Dang</collectorName>
, 
<collectorName box="[891,1091,1815,1840]" pageId="123">Ba Vuong Truong</collectorName>
, 
<collectorName box="[1102,1298,1814,1841]" pageId="123">Minh Quan Dang</collectorName>
&amp; 
<collectorName pageId="123">Nghia Son Hoang</collectorName>
, 
<collectorName box="[226,288,1851,1876]" pageId="123">Dang</collectorName>
465a
</emphasis>
(
<collectionCode box="[367,438,1850,1876]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="123" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="124" pageId="123" type="distribution">
<paragraph blockId="123.[136,1452,1778,2095]" pageId="123">
<emphasis bold="true" box="[189,345,1886,1912]" pageId="123">Distribution:</emphasis>
—Endemic to 
<collectingCountry box="[515,610,1886,1912]" name="Vietnam" pageId="123">Vietnam</collectingCountry>
(currently known only from Kon Chu Rang Nature Reserve of 
<collectingRegion country="Vietnam" name="Gia Lai" pageId="123">Gia Lai Province</collectingRegion>
).
</paragraph>
<paragraph blockId="123.[136,1452,1778,2095]" pageId="123">
<emphasis bold="true" box="[189,428,1958,1984]" pageId="123">Habitat &amp; Ecology:</emphasis>
— 
<taxonomicName authorityName="V. S. Dang, Vuong &amp; Quan" authorityYear="2022" box="[456,733,1958,1984]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="123" phylum="Tracheophyta" rank="species" species="sonlangensis">
<emphasis box="[456,733,1958,1984]" italics="true" pageId="123">Lasianthus sonlangensis</emphasis>
</taxonomicName>
occurs in the understory of the primary evergreen forest at an elevation of 
<quantity box="[276,419,1994,2021]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="123" unit="m" value="1000.0" valueMax="1100.0" valueMin="900.0">900–1100 m</quantity>
.
</paragraph>
<paragraph blockId="123.[136,1452,1778,2095]" box="[189,908,2030,2057]" pageId="123">
<emphasis bold="true" box="[189,323,2030,2056]" pageId="123">Phenology:</emphasis>
—Flowering and fruiting from March to September.
</paragraph>
<paragraph blockId="123.[136,1452,1778,2095]" box="[189,655,2066,2095]" pageId="123">
<emphasis bold="true" box="[189,405,2066,2092]" pageId="123">Vernacular name:</emphasis>
—Xú hưƠng sƠn lạng.
</paragraph>
<paragraph blockId="124.[136,1452,158,293]" pageId="124">
<emphasis bold="true" box="[189,265,158,184]" pageId="124">Notes:</emphasis>
— 
<taxonomicName authorityName="V. S. Dang, Vuong &amp; Quan" authorityYear="2022" box="[294,568,158,184]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="124" phylum="Tracheophyta" rank="species" species="sonlangensis">
<emphasis box="[294,568,158,184]" italics="true" pageId="124">Lasianthus sonlangensis</emphasis>
</taxonomicName>
was described from a Vietnamese specimen (
<emphasis box="[1078,1189,158,184]" italics="true" pageId="124">Dang 465</emphasis>
) by 
<bibRefCitation author="Dang, M. Q. &amp; Nguyen, M. H. &amp; Hoang, N. S. &amp; Le, V. T. &amp; Nguyen, T. M. H. &amp; Ho, N. Q. C. &amp; Truong, B. V. &amp; Dang, V. S." box="[1240,1448,158,185]" pageId="124" pagination="19 - 31" refId="ref53709" refString="Dang, M. Q., Nguyen, M. H., Hoang, N. S., Le, V. T., Nguyen, T. M. H., Ho, N. Q. C., Truong, B. V. &amp; Dang, V. S. (2022) Two new species of Lasianthus Jack (Rubiaceae) from southern Vietnam. European Journal of Taxonomy 806: 19 - 31. https: // doi. org / 10.5852 / ejt. 2022.806.1697" type="journal article" year="2022">
Dang 
<emphasis box="[1307,1363,158,184]" italics="true" pageId="124">et al.</emphasis>
(2022)
</bibRefCitation>
. It is similar to 
<taxonomicName authority="Pit. (1924: 390)" authorityName="Pitard" authorityPageNumber="390" authorityYear="1924" box="[307,638,194,220]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="124" phylum="Tracheophyta" rank="species" species="caeruleus">
<emphasis box="[307,448,194,220]" italics="true" pageId="124">L. caeruleus</emphasis>
Pit. (1924: 390)
</taxonomicName>
in the shape of leaves and inflorescences, but differs from the latter in having more than secondary veins, shorter petioles, bracts present (vs. absent), larger calyx and corolla lobes 4 (vs. 5).
</paragraph>
</subSubSection>
</treatment>
</document>