<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFD0FF8F5AB6F8344140CDFC" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus kbangensis V. S. Dang, Vuong &amp; Naiki 2022" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571738" ID-Zenodo-Dep="7571738" LSID="urn:lsid:plazi:treatment:43284262FFD0FF8F5AB6F8344140CDFC" httpUri="http://treatment.plazi.org/id/43284262FFD0FF8F5AB6F8344140CDFC" lastPageId="81" pageId="79">
<subSubSection box="[136,1237,1937,1964]" pageId="79" type="nomenclature">
<paragraph blockId="79.[136,1452,1937,2071]" box="[136,1237,1937,1964]" pageId="79">
<emphasis bold="true" box="[136,171,1937,1963]" pageId="79">35.</emphasis>
<taxonomicName authority="V. S. Dang, Vuong &amp; Naiki (2022: 292)" authorityName="V. S. Dang, Vuong &amp; Naiki" authorityPageNumber="292" authorityYear="2022" box="[178,882,1937,1964]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="79" phylum="Tracheophyta" rank="species" species="kbangensis">
<emphasis bold="true" box="[178,443,1937,1964]" italics="true" pageId="79">Lasianthus kbangensis</emphasis>
V.S.Dang, Vuong &amp; Naiki (2022: 292)
</taxonomicName>
— 
<figureCitation box="[912,989,1937,1964]" captionStart="FIGURE 59" captionStartId="80.[136,229,1773,1795]" captionTargetBox="[167,1418,194,1748]" captionTargetId="figure-25@80.[165,1419,193,1749]" captionTargetPageId="80" captionText="FIGURE 59. Lasianthus kbangensis V.S.Dang, Vuong &amp; Naiki (from Dang 463, VNM): A. Fruiting branch, B. Stipules, C. Abaxial leaf surfaces, D–F. Inflorescences and flower buds, G–H. Young fruits, I. Mature fruit." figureDoi="http://doi.org/10.5281/zenodo.7571734" httpUri="https://zenodo.org/record/7571734/files/figure.png" pageId="79">Fig. 59</figureCitation>
(plate), 
<figureCitation box="[1083,1162,1937,1964]" captionStart="FIGURE 57" captionStartId="78.[136,229,1796,1818]" captionTargetBox="[176,1411,204,1762]" captionTargetId="figure-24@78.[165,1418,193,1773]" captionTargetPageId="78" captionText="FIGURE 57. Distribution maps of Lasianthus inodorus, L. japonicus, L. kbangensis and L. khanhhoaensis." figureDoi="http://doi.org/10.5281/zenodo.7571728" httpUri="https://zenodo.org/record/7571728/files/figure.png" pageId="79">Fig. 57</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection lastPageId="80" pageId="79" type="materials_examined">
<paragraph blockId="79.[136,1452,1937,2071]" pageId="79">
<emphasis bold="true" box="[136,194,1977,1998]" pageId="79">Type:</emphasis>
— 
<materialsCitation collectingDate="2021-04-20" collectionCode="VNM, HN" collectorName="Van-Son Dang &amp; Ba Vuong Truong &amp; Ngoc Minh Tam Thao Duong &amp; Ngan Thi Kim Le &amp; Dinh Hiep Nguyen &amp; Dang" country="Vietnam" elevation="1015" location="Kon Chu Rang Nature Reserve" municipality="K'Bang district" pageId="79" specimenCode="VNM00045839, VNM00045840" specimenCount="1" stateProvince="Gia Lai" typeStatus="holotype">
<collectingCountry box="[219,332,1977,1998]" name="Vietnam" pageId="79">VIETNAM</collectingCountry>
. 
<collectingRegion box="[341,507,1977,1999]" country="Vietnam" name="Gia Lai" pageId="79">Gia Lai province</collectingRegion>
, 
<collectingMunicipality box="[517,668,1977,1999]" pageId="79">K’Bang district</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:43284262FFD0FF8F5AB6F8344140CDFC:CE5EA5AFFFD0FF915898F81C43F3CB86" box="[678,981,1977,1999]" country="Vietnam" municipality="K'Bang district" name="Kon Chu Rang Nature Reserve" pageId="79" stateProvince="Gia Lai">Kon Chu Rang Nature Reserve</location>
, 
<quantity box="[990,1064,1977,1999]" metricMagnitude="3" metricUnit="m" metricValue="1.015" pageId="79" unit="m" value="1015.0">
<elevation box="[990,1064,1977,1999]" metricMagnitude="3" metricUnit="m" metricValue="1.015" pageId="79" unit="m" value="1015.0">1015 m</elevation>
</quantity>
elev., 
<date box="[1126,1262,1977,1999]" pageId="79" value="2021-04-20">
<collectingDate box="[1126,1262,1977,1999]" pageId="79" value="2021-04-20">20 April 2021</collectingDate>
</date>
, 
<emphasis italics="true" pageId="79">
<collectorName box="[1273,1415,1977,1998]" pageId="79">Van-Son Dang</collectorName>
, 
<collectorName pageId="79">Ba Vuong Truong</collectorName>
, 
<collectorName box="[336,617,2013,2034]" pageId="79">Ngoc Minh Tam Thao Duong</collectorName>
, 
<collectorName box="[626,790,2013,2034]" pageId="79">Ngan Thi Kim Le</collectorName>
&amp; 
<collectorName box="[820,1001,2013,2034]" pageId="79">Dinh Hiep Nguyen</collectorName>
, 
<collectorName box="[1010,1063,2013,2034]" pageId="79">Dang</collectorName>
463
</emphasis>
(
<typeStatus box="[1117,1203,2013,2035]" pageId="79">holotype</typeStatus>
<collectionCode box="[1207,1268,2013,2034]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="79" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[1282,1442,2013,2035]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="79" type="Herbarium">VNM00045839</specimenCode>
]; isotypes 
<collectionCode box="[274,314,2049,2070]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="79" type="Herbarium">HN</collectionCode>
! 
<collectionCode box="[322,383,2049,2070]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="79" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[398,559,2049,2071]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="79" type="Herbarium">VNM00045840</specimenCode>
])
</materialsCitation>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7571734" ID-Zenodo-Dep="7571734" httpUri="https://zenodo.org/record/7571734/files/figure.png" pageId="80" startId="80.[136,229,1773,1795]" targetBox="[167,1418,194,1748]" targetPageId="80">
<paragraph blockId="80.[136,1451,1773,1832]" pageId="80">
<emphasis bold="true" box="[136,265,1773,1795]" pageId="80">FIGURE 59.</emphasis>
<taxonomicName authority="V. S. Dang, Vuong &amp; Naiki" authorityName="V. S. Dang, Vuong &amp; Naiki" authorityYear="2022" box="[272,747,1774,1796]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="80" phylum="Tracheophyta" rank="species" species="kbangensis">
<emphasis box="[272,490,1774,1795]" italics="true" pageId="80">Lasianthus kbangensis</emphasis>
V.S.Dang, Vuong &amp; Naiki
</taxonomicName>
(from 
<emphasis box="[814,909,1773,1795]" italics="true" pageId="80">Dang 463</emphasis>
, VNM): 
<emphasis bold="true" box="[998,1021,1773,1794]" pageId="80">A.</emphasis>
Fruiting branch, 
<emphasis bold="true" box="[1188,1210,1774,1795]" pageId="80">B.</emphasis>
Stipules, 
<emphasis bold="true" box="[1305,1329,1773,1795]" pageId="80">C.</emphasis>
Abaxial leaf surfaces, 
<emphasis bold="true" box="[227,274,1810,1831]" pageId="80">D–F.</emphasis>
Inflorescences and flower buds, 
<emphasis bold="true" box="[592,647,1809,1831]" pageId="80">G–H.</emphasis>
Young fruits, 
<emphasis bold="true" box="[784,799,1810,1831]" pageId="80">I.</emphasis>
Mature fruit.
</paragraph>
</caption>
<paragraph blockId="80.[136,1451,1878,2048]" pageId="80">
<emphasis bold="true" box="[136,396,1878,1904]" pageId="80">
<collectingRegion box="[136,262,1878,1904]" pageId="80">Specimens</collectingRegion>
examined:
</emphasis>
— 
<materialsCitation collectingDate="2021-04-24" collectionCode="VNM" collectorName="Van-Son Dang &amp; Ba Vuong Truong &amp; Ngoc Minh Tam Thao Duong &amp; Ngan Thi Kim Le &amp; Dinh Hiep Nguyen &amp; Dang" country="Vietnam" elevation="1010" location="Kon Chu Rang Nature Reserve" municipality="K'Bang District" pageId="80" specimenCount="1" stateProvince="Gia Lai">
<collectingCountry box="[425,557,1878,1904]" name="Vietnam" pageId="80">VIETNAM</collectingCountry>
. 
<emphasis bold="true" box="[569,670,1878,1904]" pageId="80">Central:</emphasis>
<collectingRegion box="[679,877,1878,1904]" country="Vietnam" name="Gia Lai" pageId="80">Gia Lai Province</collectingRegion>
, 
<collectingMunicipality box="[890,1075,1878,1904]" pageId="80">K’Bang District</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:43284262FFD0FF8F5AB6F8344140CDFC:CE5EA5AFFFCFFF8E5E7EF8F34581CB38" box="[1088,1447,1878,1904]" country="Vietnam" municipality="K'Bang District" name="Kon Chu Rang Nature Reserve" pageId="80" stateProvince="Gia Lai">Kon Chu Rang Nature Reserve</location>
, 
<quantity box="[136,220,1914,1940]" metricMagnitude="3" metricUnit="m" metricValue="1.01" pageId="80" unit="m" value="1010.0">
<elevation box="[136,220,1914,1940]" metricMagnitude="3" metricUnit="m" metricValue="1.01" pageId="80" unit="m" value="1010.0">1010 m</elevation>
</quantity>
elev., 
<date box="[290,446,1914,1940]" pageId="80" value="2021-04-24">
<collectingDate box="[290,446,1914,1940]" pageId="80" value="2021-04-24">24 April 2021</collectingDate>
</date>
, 
<emphasis italics="true" pageId="80">
<collectorName box="[456,620,1914,1940]" pageId="80">Van-Son Dang</collectorName>
, 
<collectorName box="[630,827,1915,1940]" pageId="80">Ba Vuong Truong</collectorName>
, 
<collectorName box="[837,1162,1914,1940]" pageId="80">Ngoc Minh Tam Thao Duong</collectorName>
, 
<collectorName box="[1172,1362,1914,1940]" pageId="80">Ngan Thi Kim Le</collectorName>
&amp; 
<collectorName pageId="80">Dinh Hiep Nguyen</collectorName>
, 
<collectorName box="[297,359,1951,1976]" pageId="80">Dang</collectorName>
463a
</emphasis>
(
<collectionCode box="[437,508,1950,1976]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="80" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="81" pageId="80" type="distribution">
<paragraph blockId="80.[136,1451,1878,2048]" pageId="80">
<emphasis bold="true" box="[189,345,1986,2012]" pageId="80">Distribution:</emphasis>
—Endemic to 
<collectingCountry box="[515,610,1986,2012]" name="Vietnam" pageId="80">Vietnam</collectingCountry>
(currently known only from Kon Chu Rang Nature Reserve of 
<collectingRegion country="Vietnam" name="Gia Lai" pageId="80">Gia Lai Province</collectingRegion>
).
</paragraph>
<paragraph blockId="81.[136,1452,158,436]" pageId="81">
<emphasis bold="true" box="[189,424,158,184]" pageId="81">Habitat &amp; Ecology:</emphasis>
— 
<taxonomicName authorityName="V. S. Dang, Vuong &amp; Naiki" authorityYear="2022" box="[452,707,158,184]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="81" phylum="Tracheophyta" rank="species" species="kbangensis">
<emphasis box="[452,707,158,184]" italics="true" pageId="81">Lasianthus kbangensis</emphasis>
</taxonomicName>
is easily found under the shade of the primary evergreen forest at an elevation of 
<quantity box="[309,466,194,221]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="81" unit="m" value="1050.0" valueMax="1100.0" valueMin="1000.0">1000–1100 m</quantity>
.
</paragraph>
<paragraph blockId="81.[136,1452,158,436]" box="[189,908,230,257]" pageId="81">
<emphasis bold="true" box="[189,323,230,256]" pageId="81">Phenology:</emphasis>
—Flowering and fruiting from March to September.
</paragraph>
<paragraph blockId="81.[136,1452,158,436]" box="[189,629,266,295]" pageId="81">
<emphasis bold="true" box="[189,405,266,292]" pageId="81">Vernacular name:</emphasis>
—Xú hưƠng kbang.
</paragraph>
<paragraph blockId="81.[136,1452,158,436]" pageId="81">
<emphasis bold="true" box="[189,265,302,328]" pageId="81">Notes:</emphasis>
— 
<taxonomicName authorityName="V. S. Dang, Vuong &amp; Naiki" authorityYear="2022" box="[294,547,302,328]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="81" phylum="Tracheophyta" rank="species" species="kbangensis">
<emphasis box="[294,547,302,328]" italics="true" pageId="81">Lasianthus kbangensis</emphasis>
</taxonomicName>
was described from a Vietnamese specimen (
<emphasis box="[1053,1164,302,328]" italics="true" pageId="81">Dang 463</emphasis>
) by 
<bibRefCitation author="Nguyen, D. H. &amp; Dang, M. Q. &amp; Le, T. K. N. &amp; Quach, V. T. E. &amp; Pham, V. N. &amp; Le, V. T. &amp; Truong, B. V. &amp; Naiki, A. &amp; Dang, V. S." box="[1213,1448,302,329]" pageId="81" pagination="291 - 296" refId="ref54877" refString="Nguyen, D. H., Dang, M. Q., Le, T. K. N., Quach, V. T. E., Pham, V. N., Le, V. T., Truong, B. V., Naiki, A. &amp; Dang, V. S. (2022) A new species of Lasianthus (Rubiaceae) from Kon Chu Rang Nature Reserve in central highlands of Vietnam. Phytotaxa 541 (3): 291 - 296. https: // doi. org / 10.11646 / phytotaxa. 541.3.9" type="journal article" year="2022">
Nguyen 
<emphasis box="[1308,1364,302,328]" italics="true" pageId="81">et al.</emphasis>
(2022)
</bibRefCitation>
. It is similar to 
<taxonomicName authority="H. Zhu (2002: 400)" authorityName="H. Zhu" authorityPageNumber="400" authorityYear="2002" box="[299,693,338,364]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="81" phylum="Tracheophyta" rank="species" species="oblongilobus">
<emphasis box="[299,475,338,364]" italics="true" pageId="81">L. oblongilobus</emphasis>
H.Zhu (2002: 400)
</taxonomicName>
in its shrub habit, smooth on branches and branchlets, and glabrous leaves, but differs in having larger leaf blades, longer petioles, calyx lobes 4 (vs. 5), smaller corolla. Fruit color is also different when ripe.
</paragraph>
</subSubSection>
</treatment>
</document>