<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFA9FFE85AB6FB2B40D9CB78" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus elevatineurus H. Zhu 2002" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571648" ID-Zenodo-Dep="7571648" LSID="urn:lsid:plazi:treatment:43284262FFA9FFE85AB6FB2B40D9CB78" httpUri="http://treatment.plazi.org/id/43284262FFA9FFE85AB6FB2B40D9CB78" pageId="54">
<subSubSection box="[136,1042,1166,1193]" pageId="54" type="nomenclature">
<paragraph blockId="54.[136,1452,1166,1264]" box="[136,1042,1166,1193]" pageId="54">
<emphasis bold="true" box="[136,171,1166,1192]" pageId="54">20.</emphasis>
<taxonomicName ID-CoL="3SCKK" authority="H. Zhu (2002: 395)" authorityName="H. Zhu" authorityPageNumber="395" authorityYear="2002" box="[178,688,1166,1193]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="54" phylum="Tracheophyta" rank="species" species="elevatineurus">
<emphasis bold="true" box="[178,470,1166,1193]" italics="true" pageId="54">Lasianthus elevatineurus</emphasis>
H.Zhu (2002: 395)
</taxonomicName>
<emphasis bold="true" box="[688,716,1166,1192]" pageId="54">—</emphasis>
<figureCitation box="[716,795,1166,1192]" captionStart="FIGURE 40" captionStartId="56.[136,229,1763,1785]" captionTargetBox="[167,1417,196,1737]" captionTargetId="figure-25@56.[166,1418,193,1739]" captionTargetPageId="56" captionText="FIGURE 40. Lasianthus elevatineurus H.Zhu (from Dang 307, VNM): A. Habit, B. Branch with fruits, C. Stipules and young fruit, D. Abaxial leaf surfaces, E–F. Inflorescence showing side view calyx and young fruits, G–I. Young and mature fruits." figureDoi="http://doi.org/10.5281/zenodo.7571672" httpUri="https://zenodo.org/record/7571672/files/figure.png" pageId="54">Fig. 40</figureCitation>
(plate), 
<figureCitation box="[889,968,1166,1192]" captionStart="FIGURE 37" captionStartId="52.[136,229,1789,1811]" captionTargetBox="[176,1405,205,1753]" captionTargetId="figure-25@52.[165,1418,193,1765]" captionTargetPageId="52" captionText="FIGURE 37. Distribution maps of Lasianthus dalatensis, L. dienkhanhensis, L. eberhartii and L. elevatineurus." figureDoi="http://doi.org/10.5281/zenodo.7571662" httpUri="https://zenodo.org/record/7571662/files/figure.png" pageId="54">Fig. 37</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection pageId="54" type="materials_examined">
<paragraph blockId="54.[136,1452,1166,1264]" pageId="54">
<emphasis bold="true" box="[136,194,1206,1227]" pageId="54">Type:</emphasis>
— 
<materialsCitation collectingDate="1997-03-22" collectionCode="AAU" collectorName="Averyanov L. &amp; Nguyen Q. B. &amp; Phan K. L." country="Vietnam" elevation="2150" location="Lac Duong District" pageId="54" specimenCount="1" stateProvince="Lam Dong" typeStatus="holotype">
<collectingCountry box="[219,332,1206,1227]" name="Vietnam" pageId="54">VIETNAM</collectingCountry>
. 
<collectingRegion box="[343,543,1206,1228]" country="Vietnam" name="Lam Dong" pageId="54">Lam Dong Province</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:43284262FFA9FFE85AB6FB2B40D9CB78:CE5EA5AFFFA9FFE85814FB1342CCC884" box="[554,746,1206,1228]" country="Vietnam" name="Lac Duong District" pageId="54" stateProvince="Lam Dong">Lac Duong District</location>
, 
<quantity box="[756,891,1206,1227]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="54" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">
<elevation box="[756,891,1206,1227]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="54" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">2100–2200 m</elevation>
</quantity>
elev., 
<date box="[957,1108,1206,1228]" pageId="54" value="1997-03-22">
<collectingDate box="[957,1108,1206,1228]" pageId="54" value="1997-03-22">22 March 1997</collectingDate>
</date>
, 
<emphasis italics="true" pageId="54">
<collectorName box="[1120,1251,1206,1227]" pageId="54">Averyanov L.</collectorName>
, 
<collectorName box="[1262,1389,1206,1227]" pageId="54">Nguyen Q.B.</collectorName>
, 
<collectorName pageId="54">Phan K.L.</collectorName>
VH 3045
</emphasis>
(
<typeStatus box="[335,421,1242,1264]" pageId="54">holotype</typeStatus>
<collectionCode box="[425,486,1242,1263]" country="Ethiopia" name="Addis Ababa University, Department of Biology" pageId="54">AAU</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<paragraph blockId="54.[136,1452,1310,1840]" pageId="54">
<emphasis bold="true" box="[136,396,1310,1336]" pageId="54">
<collectingRegion box="[136,262,1310,1336]" pageId="54">Specimens</collectingRegion>
examined:
</emphasis>
— 
<materialsCitation collectingDate="2019-05-09" collectingDateMax="2019-06-11" collectingDateMin="2019-05-09" collectionCode="VNM" collectorName="Dang V. S. &amp; Hoang N. S. &amp; Dang" country="Vietnam" county="Gia Nghia District" elevation="810" location="Ta Dung National Park" municipality="Dang" pageId="54" specimenCount="1" stateProvince="Dak Nong">
<collectingCountry box="[425,557,1310,1336]" name="Vietnam" pageId="54">VIETNAM</collectingCountry>
. 
<emphasis bold="true" box="[570,671,1310,1336]" pageId="54">Central:</emphasis>
<collectingRegion box="[680,910,1310,1337]" country="Vietnam" name="Dak Nong" pageId="54">Dak Nong Province</collectingRegion>
, 
<collectingCounty box="[924,1139,1310,1336]" pageId="54">Gia Nghia District</collectingCounty>
, 
<quantity box="[1153,1226,1310,1337]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="54" unit="m" value="810.0">
<elevation box="[1153,1226,1310,1337]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="54" unit="m" value="810.0">810 m</elevation>
</quantity>
elev., 
<date box="[1304,1447,1310,1337]" pageId="54" value="2019-05-09">
<collectingDate box="[1304,1447,1310,1337]" pageId="54" value="2019-05-09">9 May 2019</collectingDate>
</date>
, 
<emphasis box="[136,537,1346,1373]" italics="true" pageId="54">
<collectorName box="[136,247,1347,1373]" pageId="54">Dang V.S.</collectorName>
&amp; 
<collectorName box="[282,415,1347,1373]" pageId="54">Hoang N.S.</collectorName>
, 
<collectingMunicipality box="[426,488,1347,1372]" pageId="54">Dang</collectingMunicipality>
307
</emphasis>
(
<collectionCode box="[552,623,1346,1372]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="54" type="Herbarium">VNM</collectionCode>
); in the same location, 
<location LSID="urn:lsid:plazi:treatment:43284262FFA9FFE85AB6FB2B40D9CB78:CE5EA5AFFFA9FFE8594FFAE64450C914" box="[881,1142,1346,1373]" country="Vietnam" county="Gia Nghia District" municipality="Dang" name="Ta Dung National Park" pageId="54" stateProvince="Dak Nong">Ta Dung National Park</location>
, 
<quantity box="[1153,1224,1346,1373]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="54" unit="m" value="680.0">
<elevation box="[1153,1224,1346,1373]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="54" unit="m" value="680.0">680 m</elevation>
</quantity>
elev., 
<date box="[1296,1447,1346,1373]" pageId="54" value="2019-06-11">
<collectingDate box="[1296,1447,1346,1373]" pageId="54" value="2019-06-11">11 June 2019</collectingDate>
</date>
, 
<emphasis box="[136,538,1382,1409]" italics="true" pageId="54">
<collectorName box="[136,247,1383,1409]" pageId="54">Dang V.S.</collectorName>
&amp; 
<collectorName box="[283,415,1383,1409]" pageId="54">Hoang N.S.</collectorName>
, 
<collectorName box="[426,488,1383,1408]" pageId="54">Dang</collectorName>
309
</emphasis>
(
<collectionCode box="[553,624,1382,1408]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="54" type="Herbarium">VNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="54" type="distribution">
<paragraph blockId="54.[136,1452,1310,1840]" pageId="54">
<emphasis bold="true" box="[189,345,1418,1444]" pageId="54">Distribution:</emphasis>
—Endemic to 
<collectingCountry box="[509,604,1418,1444]" name="Vietnam" pageId="54">Vietnam</collectingCountry>
(currently known as Gia Nghia District and Ta Nung National Park of 
<collectingRegion country="Vietnam" name="Dak Nong" pageId="54">Dak Nong Province</collectingRegion>
, and Lac Duong District and Bidoup Nui Ba National Park of 
<collectingRegion box="[1006,1238,1454,1481]" country="Vietnam" name="Lam Dong" pageId="54">Lam Dong Province</collectingRegion>
).
</paragraph>
<paragraph blockId="54.[136,1452,1310,1840]" pageId="54">
<emphasis bold="true" box="[189,426,1490,1516]" pageId="54">Habitat &amp; Ecology:</emphasis>
—Growing in primary and secondary montane forests, edge of evergreen forest with open shaded area, at 
<quantity box="[309,450,1526,1553]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="2.2" metricValueMin="0.6" pageId="54" unit="m" value="1400.0" valueMax="2200.0" valueMin="600.0">600–2200 m</quantity>
elev.
</paragraph>
<paragraph blockId="54.[136,1452,1310,1840]" box="[189,1059,1562,1589]" pageId="54">
<emphasis bold="true" box="[189,323,1562,1588]" pageId="54">Phenology:</emphasis>
—Flowering and fruiting from December to July of the next year.
</paragraph>
<paragraph blockId="54.[136,1452,1310,1840]" box="[189,637,1598,1627]" pageId="54">
<emphasis bold="true" box="[189,405,1598,1624]" pageId="54">Vernacular name:</emphasis>
—Xú hưƠng gân lồi.
</paragraph>
<paragraph blockId="54.[136,1452,1310,1840]" pageId="54">
<emphasis bold="true" box="[189,265,1634,1660]" pageId="54">Notes:</emphasis>
— 
<taxonomicName box="[294,575,1634,1660]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="54" phylum="Tracheophyta" rank="species" species="elevatineurus">
<emphasis box="[294,575,1634,1660]" italics="true" pageId="54">Lasianthus elevatineurus</emphasis>
</taxonomicName>
was reported as new to 
<collectingCountry box="[850,945,1634,1660]" name="Vietnam" pageId="54">Vietnam</collectingCountry>
by Zhu (2002). This species is characterized by having an elliptic-lanceolate leaf blade, secondary veins 7–9 on each side of midrib, bracts linear, hirsute, calyx obconical, and fruits globose, white when ripe.
</paragraph>
<paragraph blockId="54.[136,1452,1310,1840]" pageId="54">
This species is morphologically similar to 
<taxonomicName authority="Blume (1826: 997)" authorityName="Blume" authorityPageNumber="997" authorityYear="1826" box="[668,1000,1742,1768]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="54" phylum="Tracheophyta" rank="species" species="lucidus">
<emphasis box="[668,779,1742,1768]" italics="true" pageId="54">L. lucidus</emphasis>
Blume (1826: 997)
</taxonomicName>
based on its shape of stipules and calyx but differs from the latter in having 7–9 (vs. 3–6) secondary veins on each side of midrib and white (vs. blue) fruits when ripe.
</paragraph>
</subSubSection>
</treatment>
</document>