Resurrection of Stipa tremula and taxonomy of the high-alpine species from the Stipa purpurea complex (Poaceae, Pooideae) Author Nobis, Marcin https://orcid.org/0000-0002-1594-2418 Institute of Botany, Jagiellonian University, Gronostajowa 3, 30 - 387 Krakow, Poland m.nobis@uj.edu.pl Author Krzempek, Marta Institute of Botany, Jagiellonian University, Gronostajowa 3, 30 - 387 Krakow, Poland Author Nowak, Arkadiusz https://orcid.org/0000-0001-8638-0208 Institute of Biology, University of Opole, 45 - 052 Opole, Poland & Botanical Garden, Center for Biological Diversity Conservation, Polish Academy of Sciences, 02 - 976 Warszawa, Poland Author Gudkova, Polina D. https://orcid.org/0000-0002-6537-8018 Research Laboratory ' Herbarium', National Research Tomsk State University, Lenin 36 Ave., 634050 Tomsk, Russia & Institute of Biology, Altai State University, Lenin 61 Ave., 656049, Barnaul, Russia Author Klichowska, Ewelina https://orcid.org/0000-0001-9641-5750 Institute of Botany, Jagiellonian University, Gronostajowa 3, 30 - 387 Krakow, Poland ewelina.klichowska@uj.edu.pl text PhytoKeys 2022 2022-05-13 196 21 47 http://dx.doi.org/10.3897/phytokeys.196.82598 journal article http://dx.doi.org/10.3897/phytokeys.196.82598 1314-2003-196-21 A221AFB3297756818D82E7B978F2E682 Stipa x ladakhensis M. Nobis, Klichowska, A. Nowak & P.D. Gudkova, nothosp. nov.(S. klimesii x S. purpurea s.l.) Fig. 5 Type . NW India , Jammu and Kashmir State , Ladakh Region : Zanskar : Zara , Spangchen Do , alt. 4520 m , 1 Sep 2001 , 33°22.7'N , 77°45.1'E , code 01-34-13, L. Klimes 1474 ( holotype KRA 603490!, isotypes PR!, KRA 603487!, 603486!) Figure 5. The holotype of Stipa x ladakhensis M. Nobis, Klichowska, A. Nowak & P.D. Gudkova. Description. Plant perennial , densely tufted, with a few culms and numerous vegetative shoots; culms 35-55 cm tall, 1-2-noded, nodes distributed close together and only in the lowermost part of the culm. Leaves of vegetative shoots : sheaths shortly and densely pubescent; ligules acute, on the external sheaths (1.0-)1.2-2.0(-2.8) mm long, whereas, on the internal sheaths (1.5-)2.0-4.0(-5.5) mm long; blades convolute, green, pale green to glaucous 10-25 cm long, 0.3-0.5 mm in diameter, adaxial surface covered by 0.15-0.2 mm long hairs, abaxial surface scabrous. Cauline leaves : lower sheaths shortly pubescent, upper scabrous or glabrous; ligules acute, (1.0-)2.0-4.0(-5.0) mm long; blades of convolute, green or pale green, adaxial surface shortly pubescent, abaxial surface scabrous or glabrous. Panicle 17-25 cm long, rather contracted, with 15-23 spikelets, at base enclosed by the sheath of the uppermost leaf or exerted, lower branches 2-6 cm long, straight or slightly flexuous, setulose or glabrous, single or paired. Glumes subequal, brownish to purplish, glumes 16-19(-20) mm long, narrowly lanceolate, tapering into long hyaline apex. Floret (lemma + callus) 8.5-10.2 mm long and 0.7-0.9 mm wide. Callus 1.5-1.9 mm long, densely and long-pilose, the base of callus narrow, peripheral ring 0.15-0.20 mm in diameter, acute, scar narrow elliptic. Lemma coriaceous, straw-coloured, brownish or purplish; covered throughout (from the bottom to top) by dense ascending to appressed hairs 0.2-0.4 mm long. Awn 46-58 mm long, bigeniculate, lower segment of the awn 8-11 mm long, twisted, with 1.7-2.3 mm long hairs, middle segment of the awn 4-9 mm long, twisted, with 1.6-2.2 mm long hairs; terminal segment of the awn (seta) flexuous 30-42 mm long with hairs shorter to equal to those on the column, 1.6-2.1 mm long, gradually decreasing in length towards the apex. Palea equalling lemma in length. Habitat. High mountain semi-deserts, on the elevation from 4000 to 5000 m. Distribution. W Himalayas (NW India).