Taxonomic notes on Hydroidomedusae (Cnidaria) from South China Sea III: Family Rathkeidae and Zancleopsidae
Author
Wang, Chunguang
Third Institute of Oceanography, State Oceanic Administration, Xiamen 361005, China
Author
Xu, Zhenzu
College of Ocean and Earth Sciences, Xiamen University, Xiamen 361102, China
Author
Huang, Jiaqi
College of Ocean and Earth Sciences, Xiamen University, Xiamen 361102, China
Author
Guo, Donghui
College of Ocean and Earth Sciences, Xiamen University, Xiamen 361102, China
Author
Lin, Mao
Third Institute of Oceanography, State Oceanic Administration, Xiamen 361005, China & Collaborative Innovation Center of Deep Sea Biology, Hangzhou 310012, China
Author
Xia, Zhen
Guangzhou Marine Geological Survey, China Geological Survey, Guangzhou 510760, China Corresponding author, E-mail: guodh @ xmu. edu. cn
text
Zoological Systematics
2016
41
4
392
403
journal article
10.11865/zs.201644
2095-6827
5366513
DDA913E3-9578-476B-AFFB-E6C5392C430D
Zancleopsis oblongus
Xu, Huang & Wang
,
sp. nov.
(
Figs 16–21
)
Material
examined.
Holotype
(
TIO 022
),
South
China
Sea
, station NNXW12095 (
11°58'N
,
113°02'E
), depth 4294 –
230 m
,
1 September 2012
, coll.
Peng Xiang
(
TIO
).
Description. Mesuda up to 3.5 mm high, with apical projection, reach one-third of the total height, umbrella almost conical-shaped, higher than width, jelly thick, forming pointed apex; manubrium flask-shaped, about half as long as the bell cavity, with large quadratic base, distal part of manubrium simple, conical, mouth opening quadrangular with faint lips, margin thick provided with cnidocysts; gonads interradial, with shallow groove which may divide them into 8 adradial masses, but separated perradially; 4 radial canals and a ring canal; 2 long, opposite capitate tentacles, each with 60–65 buttony-shaped cnidocysts knobs along the whole tentacle on the abaxial side, and 2 shorter, opposed capitate tentacles, simple, terminating in a slight swelling; 4 marginal tentacular bulbs clasping umbrella margin, of which 2 long tentacular bulbs oblong-shaped, without adaxial expansion at the base of the tentacles, and other 2 shorter tentacular bulbs very small, conical-shaped, with a large, spherical-shaped adaxial expansion at tentacular base covered with cnidocysts; with abaxial ocelli on marginal bulbs; velum narrow.
Biology. A deep-water species, occurring at depths of 4294–
230 m
.
Distribution. This species is rare in the southern of South
China
Sea, it is only known from
one specimen
caught in deep waters.
Etymology. The species name is from the Latin
oblongus
, meaning oblong, referring to the morphology of two opposite long tentacular bulbs.
Remarks. This new species has following characters which are in common with
Zancleopsis
Hartlaub, 1907
: umbrella with apical projection; two long and short tentacles; marginal bulbs with two adaxial cnidocysts knobs; gonads on interradial regional of manubrium with interradial grooves; abaxial ocelli on the marginal bulbs.
The new species is similar to
Z. tentaculata
Kramp, 1928
, but differs from the latter by following: 1) manubrium quadratic base and mouth square; 2) two long opposite tentacles with 60–65 buttony-shaped cnidocysts knobs along the whole tentacles; 3) two long tentacular bulbs oblong-like, without adaxial cnidocysts knobs; 4) gonads interradial with shallow grooves.
Figures 16–18.
Zancleopsis oblongus
Xu, Huang & Wang
,
sp. nov.
16. Lateral view. 17. Enlarged lateral view of long tentacular bulb. 18. Enlarged lateral view of short tentacular bulb. Scale bars: 16 =1.0 mm; 17–18 = 0.1 mm.
Key to medusa of all known
Zancleopsis
species.
1. Without apical projection; two long tentacles with 8–10 capitate side branches; four large hemispherical adacial cnidocysts knobs at tentacular base..............................................................................................................................................
Z. elegans
Bouillon, 1978
With
apical projection .......................................................................................................................................................................... 2
2. Four long capitate tentacles, each distal end with 7–15 knob-shaped cnidocysts...................................
Z. symmetrica
Bouillon, 1985
Two
long or four capitate tentacles ......................................................................................................................................................3
3. Four short capitate, club-shaped tentacles and four large cnidocysts knobs at tentacular base ........................
Z. gotoi
(
Uchida, 1927
)
Two
long capitate tentacles .................................................................................................................................................................. 4
4. Two long capitate tentacles with 2–4 capitate side branches and two opposed rudimentary bulbs .........
Z. dichotoma
(
Mayer, 1900
)
*
Two long and two short capitate tentacles............................................................................................................................................5
5. Each long tentacle with 6–15 capitate side branches at distal end of tentacles; four large, adaxial cnidocysts knobs at all tentacular base ..........................................................................................................................................................
Z. tentaculata
Kramp, 1928
Each
long tentacle with 60–65 buttony-shpaed abaxial cnidocysts knobs along whole tentacles; only two large, adaxial cnidocysts knobs at short tentacular bulbs; long tentacular bulbs oblong-shaped, without adaxial cnidocysts knobs ............................................. ........................................................................................................................................
Z. oblongus
Xu, Huang & Wang
,
sp. nov.
*
*
Presented in
China
Sea.
Funding
This study was supported by National Natural Science Foundation of
China
(41406216), Global Climate Change and Ocean Atmosphere Interaction Research (GASI-01-02-02-03, GASI-01-02-04, GASI-01-02-02-01), National Special Project on Gas Hydrate (GZH201100311), the Marine Biological Sample Museum of the Chinese Offshore Investigation and Assessment, and the National Basic Research Program of
China
(2011CB403604).