World revision of the wasp genus Liosphex Townes, 1977 (Hymenoptera: Rhopalosomatidae) Author Lohrmann, Volker Author Ohl, Michael text Zootaxa 2010 2384 1 43 journal article 10.5281/zenodo.193731 4bf51a1d-077f-46dc-bd30-e582ffbf11ce 1175-5326 193731 Liosphex darien Lohrmann sp. nov. ( Figs 38–40 ) Holotype . Female. PANAMA : Darién , Parque Nacional Darién , Estac. Rancho Frio, Pirre, 3–17 April 2002 , R.C., A.S. & R.M., Malaise trap ( MIUP ). Paratypes . PANAMA : same data as holotype , but 16 November 2000 – 17 January 2001 , 80m, R. Cambra & A. Santos (2 Ƥ, MIUP ); same data as holotype , but 11–16 December 2004 , R. Cambra, Malaise trap (1 Ƥ, MIUP ); Panama , Metropolitan, Malaise trap, ANAE 2, 6–10 Mayo 1994 , V. Rodriguez (1 3, MIUP ); same data, but 10–17 May 1994 (1 3, MIUP ). Diagnosis. This species is unique among all Liosphex in showing the combination of the following characters: UID 1.7–1.8× as long as LID (a character which this species shares only with L. micropterus , all other New World species show a UID:LID lower than 1.6) and pale markings on the base of tergites II and III (the metasoma is completely black in L. micropterus ). Description of the female. Head ( Fig. 38 ): Maximum width in frontal view 1.7 mm [ 1.6–1.7 mm ]. UID 1.7–1.8× LID. Distance between eye and torulus about same width as distance between eye and clypeus. Flagellomeres short (flagellomere II 3.1× [3.1–3.2×], flagellomere VII is 1.9× [1.9–2.1×] as long as wide). Apical bristles present on flagellomeres I–V [I–VI], longer bristle on flagellomere I 0.95× its length. OOD 2.3× [2.3–2.5×], IOD 0.9× [0.9–1.0×], MOD 1.1× [1.0–1.1×] LOD. Mesosoma: Length of mesosoma: 2.8 mm [ 2.7–2.8 mm ]. Anterior margin of pronotum without median notch (as in Fig. 20 ). FIGURES 38–40. Liosphex darien sp. nov. (holotype) Ƥ. 38: head in dorsal view; 39: habitus in lateral view; 40: habitus in dorsal view. Wings: Maximum length of forewing: 6.5 mm [ 5.8–6.5 mm ], 2.2–2.3× as long as mesosoma. Forewing 1cu-a almost vertical, distad of M by 2.05× [2.05–2.35×] its length. M-cu distinctly curved. 1Rs 2.05–2.35× as long as high, its basal angle smaller than 90° ( Fig. 9 ). Legs: Forebasitarsomere short, length of antennal cleaner 0.45× [0.4–0.45×] its length. Mid- and hindfemur with a few bristles spread over its distal dorsal area. Hindtibia with two bristles on only one side of insertion of shorter apical spur (as in Fig. 22 ). Metasoma: Tergite I 1.8–1.9× as long as wide. Color (as in Figs 39–40 ): Head and mesosoma basically dark brown to black except the following: Mouthparts except mandibular teeth and maxillar palpus, labrum, clypeus, gena, inner part of eye notch, anterior and posterior margin of pronotum, tegula, posterior end of scutellum and median section of metanotum whitish. Foreleg pale yellow to whitish with brown markings, tarsi brown. Mid- and hindleg dark brown with yellow markings on midcoxa and the base of mid and hindtibia, tarsi dark brown. Dorsolateral side of antennae brown, ventrolateral side yellow to orange. Metasoma dark brown except the following: posterior end of tergite I, anterior section of tergites and sternites II–III pale yellow to whitish. Forewing slightly brownish, more intensively at its distal section. Pilosity: Body, including legs, largely covered with dense setae. Setae lacking on following structures: occiput, ventrolateral side of pronotum, ventrolateral side of scutellum, median area of mesepisternum, ventral part of metepimeron, anteroventral of metepisternum, median part of lateral side of propodeum and posteroventral half of hindfemur. Description of the male. Head, mesosoma and metasoma as in female except the following: Head: Maximum width in frontal view 1.2 mm (approximated since the heads of both males are deformed). UID 1.7–1.75× LID. Flagellomere II 2.7–2.8×, flagellomere VII 2.0–2.2× as long as wide. IOD 1.1–1.25× LOD. Mesosoma: Length of mesosoma: 2.1 mm (approximated since the anterior margin of mesosoma hidden by head). Wings: Maximum length of forewing: 4.7 mm , 2.2× as long as mesosoma. Legs: Tarsomeres II–IV with plantulae. Life history. Almost nothing is known about the life history of Liosphex darien . Only four females and two males are known, and these were collected from mid November to May. All specimens were collected with Malaise traps. Etymological note. The species name refers to the name of the province and national park (‘ Darién ’) where the female type specimens were collected; it is a noun in apposition.