Haasea Verhoeff, 1895 - a genus of tumultuous history and chaotic recordsredefinition, revision of taxonomy and geographic distributions, with descriptions of two new species from Austria and Serbia (Diplopoda, Chordeumatida, Haaseidae) Author Antić, Dragan Author Akkari, Nesrine text Zootaxa 2020 2020-06-16 4798 1 1 77 journal article 21683 10.11646/zootaxa.4798.1.1 d8c0dac0-6b03-43f4-be6d-1dc3d3d9cfa3 1175-5326 3896551 4830ED2F-DB55-468E-97D4-2A278CBA6CE0 Haasea plasana ( Verhoeff, 1899 ) Figs 22 , 23 , 38 , 42 Orobainosoma (Brachybainosoma) plasanum Verhoeff, 1899 Orobainosoma flavescens — Attems (1929) (misidentification) Diagnosis. Differs from the other congeners, with exception of H. intermedia , H. lacusnigri , H. microcorna and H. makarovi sp. nov. , by the presence of strongly developed anterior lateral process on angioxocite. Clearly differs from those species by the presence of distally bifurcated anterior lateral process (vs. simple anterior lateral process in H. intermedia , H. lacusnigri , H. microcorna and H. makarovi sp. nov. ). Type material examined: NHMW: ♂ , Lectotype , here designated, Plasa near Jablanica , Bosnia and Herzegov- ina, Verhoeff leg. ( NHMW9824 ). Paralectotype : 1 ♀ , same data as lectotype ( NHMW2130 ). Paralectotypes ZMB: 1 juvenile ZMB3282 [ Herzegovina ]; ZMB12760 (a: anterior gonopods, posterior gonopods, lp 10, lp 11; b: 1 pair of posterior gonopods, 3 pairs of anterior gonopods, c: anterior gonopods, posterior gonopods [no precise locality], d: posterior gonopod, lp 10, 4 pair of legs (2 with tarsal papillae) [ Trebević near Sarajevo ], e: anterior gonopods probably part of slide d; ZMB12761: anterior gonopods, posterior gonopods, lp 10, lp 11 [ Trebević near Sarajevo ]; ZMB12762: anterior body section [ Plasa , Herzegovina ]. FIGURE 21. Haasea musimontium ( Strasser, 1937 ) . A. Paralectotype female (MHNG), habitus, lateral view. B–E. Lectotype male (MHNG). B. Microslides with dissected parts. C. Posterior gonopods, anterior view. D. Left anterior gonopod, mesal view. E. Right anterior gonopod, lateral view. F. Lectotype male and paralectotype female (MHNG), habitus. Abbreviations: a —angiocoxite; ab —anterior branch; dp —distal part; mp —mesal process; pb —posterior branch; p c—papillated cone; pl —posterior lamella; plp —posterior lateral process; t —telopodite. Without scale. Photos credit: Peter Schwendinger. FIGURE 22. Haasea plasana ( Verhoeff, 1899 ) . A. Male from Trebević, Bosnia & Herzegovina, habitus, lateral view. B. Male from Kopaonik Mt., Serbia, right anterior gonopod, mesal view. C. Male from Kopaonik Mt., Serbia, right anterior gonopod, lateral view. Abbreviations: alp —anterior lateral process; mp —mesal process; pc —papillated cone; pl —posterior lamella; plp —posterior lateral process; sch —seminal chamber; so —seminal opening. Scale bars: 1 mm ( A ), 0.1 mm ( B, C ). FIGURE 23 . Haasea plasana ( Verhoeff, 1899 ) , males from Kopaonik Mt., Serbia. A. Left anterior gonopod, lateral view. B. Left anterior gonopod, mesal view. C. Left anterior gonopod, detail of distal part, lateral view. D. Left anterior gonopod, detail of distal part, mesal view. E. Left posterior gonopod, mesal view. F. Posterior gonopods, anterior view. Abbreviations: a —angiocoxite; ab —anterior branch; alp —anterior lateral process; asl —anterior sternal lobe; cp —coxal process; dp —distal part; mb —medial branch; mdl —mesodistal lobe; ml —“ Opuntia -like” membranous lobe; mp —mesal process; pb —posterior branch; p c—papillated cone; pl —posterior lamella; plp —posterior lateral process; so —seminal opening; t —telopodite. Scale bars: 0.1 mm ( A , B , E , F ), 0.05 mm ( C , D ). Other material examined: BOSNIA & HERZEGOVINA : Trebević: 4 ♀♀ , 2 ♂♂ (one photographed), Brachybainosoma plasanum Attems det. ( NHMW9819 ) , 1 ♂ anterior body section, 6 posterior body sections, 2 heads, 1 mid-body section, 1 ♂ with gonopods dissected, gonopods in tube, same locality, Brachybainosoma plasanum Attems det. ( NHMW9821 ) ; 1 dissected ♂ with gonopods in tube, same locality, Brachybainosoma plasanum , Attems det. ( NHMW9820 ) . SERBIA : 3 ♂♂ (in pieces, one used for SEM), S. Makarov det., Petrova Ravan , Ravnište , Mt. Kopaonik , pit-fall traps 20.08.2002 – 23.07.2003 ., D. Pavićević leg. ( IZB ) , 1 ♂ microslide (body and gonopods), same data as previous ( IZB ) . Distribution ( Figs 38 , 42 ). Bosnia & Herzegovina : Plasa Mt., near Jablanica ( Verhoeff 1899 ), Trebević Mt., near Sarajevo ( Verhoeff 1899 ). Serbia : Petrova Ravan, Ravnište, Mt. Kopaonik ( Makarov et al. 2004 ). Type locality. Plasa Mt. in Bosnia & Herzegovina . Comments. The distance between Serbian and Bosnian localities is some 200 km . Given that western and southwestern Serbia , as well as Bosnia and Herzegovina , are very poorly studied in terms of millipede fauna, this gap will certainly be filled in the future. Evidence for this is that some millipedes described from the same localities in Bosnia and Herzegovina , are recently found in the territory of southwestern Serbia . Attems (1929: 318) stated the locality Trebević as the new locality for Orobainosoma flavescens even if, just a few sections below, he stated both Verhoeff’s (1899) localities—Plasa and Trebević for Brachybainosoma plasanum . From Attems catalogue in NHMW ( Fig. 10 ) for Craspedosoma / Orobainosoma flavescens the locality “Trebević” is stated under the number 17. Later, Attems crossed out the name of this locality for O. flavescens and wrote Brachybainosoma next to “Trebević”. It is obvious that Attems recognized the mistake and autocorrected himself. Later, he stated Brachybainosoma plasanum in the vial with specimens from Trebević.