A new species of Cardepia Hampson, 1905 (Lepidoptera: Noctuidae) from China
Author
Gyulai, Peter
H- 3530 Miskolc, Mélyvölgy 13 / A, Hungary. E-mail: gyulainegarai. adrienne @ upcmail. hu
Author
Saldaitis, Aidas
Nature Research Centre, Akademijos 2, LT- 08412 Vilnius, Lithuania. E-mail: saldrasa @ gmail. com
text
Zootaxa
2014
2014-06-23
3821
3
391
397
journal article
5341
10.11646/zootaxa.3821.3.8
beca05a3-4857-4f22-8686-b73b0d159099
1175-5326
4920121
DEB33D0C-0BC6-4189-86C6-EA54537204E9
Cardepia rothei
Gyulai & Saldaitis
sp. n.
(
Figs 1–3
,
14–17
,
30
)
Type material.
Holotype
: male (
Fig. 1
),
China
,
Xinjiang
,
W
Taklimakan
desert,
Yarkan He
riv. valley, tugay forest,
1140 m
,
N 39°21.953
’’,
E 078°11.639
’’,
9–12.
VI
. 2013
, leg.
A
. Floriani
, coll.
P
. Gyulai
(to be deposited in the
Hungarian Natural History Museum
,
Budapest
,
Hungary
); (
Slide No.
PGY
3680m
)
Paratypes
:
102 males
(
Fig. 2
),
43 females
(
Fig. 3
), with the same data as the
holotype
;
11 males
,
3 females
,
China
,
Xinjiang
, SW from Kashi,
W
Taklimakan
desert, Terambasar,
1200 m
,
N 39°10.564
’’,
E 077°04.039
’’,
7.
VI
. 2013
, leg.
A
. Floriani
, in the collections of
A
. Floriani
(
Milan
,
Italy
),
P
. Gyulai
(
Miskolc
,
Hungary
),
F
. Hofer
(
Baden
,
Austria
),
Nature Research Centre
(
Vilnius
,
Lithuania
),
D. Nilsson
(
Kalvehave
,
Denmark
),
S
. Rothe
(
Taucha
,
Germany
),
H
. Seibald
(
Vienna
,
Austria
) and
World Insect Gallery
(
Joniškis
,
Lithuania
).
Slide Nos
: PGY 3610, 3672, 3677 (males), 3616, 3738 (females)
.
Diagnosis and description.
Wingspan
23–27 mm
. Although
Cardepia rothei
sp. n.
resembles all known Central Asiatic species and subspecies of
Cardepia
it can be separated externally by its more elongate forewing apex, paler forewing ground color with less brown or gray suffused wings, and fawn or pale ochre body hair.
Cardepia rothei
is most similar to the light coloured
C. helix helix
Boursin, 1962
(
Figs 4, 5
) but is easily distinguished by its even, unserrated, and slightly wavy subterminal line which is not as medially projected toward the margin as in
C. helix
. On average
Cardepia rothei
is smaller than
C. helix
(
24-31 mm
), and the most geographically widely distributed
C. irrisoria
(Ershov, 1874)
(=
albipicta
Christoph, 1884 (
25-34 mm
) (
Figs11, 12
).
C. rothei
(
Figs 1–3
) lacks the broad white antemedial and postmedial transverse line shading of
C. kaszabi
Sukhareva & Varga, 1973
, and the white-filled orbicular spot and white -variegated forewing cilia of
C. irrisoria
f.
albipicta.
The claviform stigma is small but more conspicuous than that of
C. irrisoria
. The undersides of the wings, whose ground colors are pale fawn or whitish with subtle brown suffusion restricted to the submarginal area or to the forewing postmedial and hindwing medial lines, distinguish the new species from its Central Asian congeners except for light forms of
C. helix
and
C. irrisoria
. The underside characters are useful for separating
C. rothei
from
C. helix dubatolovi
Hacker, 1998
(
Figs 7, 6
) and
C. dardistana
Boursin, 1967
(
Figs 9, 10
). The latter species appears to be the most closely allied to
C. rothei
species based on genitalia but differs in having a distinctive and diffuse brown stripe in the ventral submarginal area.
FIGURES 1–8.
Cardepia
ssp. adults
. 1.
C. rothei
sp. n.
, male, holotype, China, Xinjiang (coll. P. Gyulai); 2.
C. rothei
sp. n.
, male, paratype, China, Xinjiang (coll. P. Gyulai); 3.
C. rothei
sp. n.
, female, paratype, China, Xinjiang (coll. P. Gyulai); 4.
C. helix helix
,
male, Turkmenistan, Mari (coll. P. Gyulai); 5.
C. helix helix
,
female, Turkmenistan, Mari (coll. P. Gyulai); 6.
C. helix dubatolovi
,
male, Tadjikistan, Tigrovaya Balka (coll. P. Gyulai); 7.
C. helix dubatolovi
,
female, Tadjikistan, Tigrovaya Balka (coll. P. Gyulai); 8.
C. irrisoria nigrescens
male, holotype, Mongolia, Bayankhongor Aimak (coll. P. Gyulai).
FIGURES 9–13.
Cardepia
ssp. adults
. 9.
C. dardistana
,
male, Pakistan, Teru (coll. P. Gyulai); 10.
C. dardistana
,
female, Pakistan, Gupis (coll. P. Gyulai); 11.
C. irrisoria irrisoria
,
male, Uzbekistan, Bukhara (coll. P. Gyulai); 12.
C. irrisoria irrisoria
,
female, Kazakhstan, Batkul lake (coll. P. Gyulai); 13. Desert river floodplain leafy forest (tugay) – the biotope of
C. rothei
sp. n.
, China, Xinjiang, Yarkan He riv. valley.
FIGURES 14–21.
Cardepia
ssp. male
genitalia. 14.
C. rothei
sp. n.
,
holotype, capsule, prep. PGy3680; 15.
C. rothei
sp. n.
, holotype, aedeagus, prep. PGy3680; 16.
C. rothei
sp. n.
,
paratype, capsule, prep. PGy3677; 17.
C. rothei
sp. n.
, paratype, aedeagus, prep. PGy3677; 18.
C. dardistana
, capsule, prep. PGy3701; 19.
C. dardistana
aedeagus, prep. PGy3701; 20.
C. helix helix
, capsule, prep. PGy3757; 21.
C. helix helix
aedeagus, prep. PGy3757.
FIGURES 22–29.
Cardepia
ssp. male
genitalia. 22.
C. helix dubatolovi
, capsule, prep. PGy2108; 23.
C. helix dubatolovi
aedeagus, prep. PGy2108; 24.
C. irrisoria irrisoria
, capsule, Uzbekistan, prep. PGy3760; 25.
C. irrisoria irrisoria
,
aedeagus, Uzbekistan, prep. PGy3760; 26.
C. irrisoria irrisoria
, capsule, China, Xinjiang, prep. PGy3696; 27.
C. irrisoria irrisoria
aedeagus, China, Xinjiang, prep. PGy3696; 28.
C. irrisoria nigrescens
, holotype, capsule, prep. HH10565; 29.
C. irrisoria nigrescens
,
holotype, aedeagus, prep. HH10565.
FIGURES 30–33.
Cardepia
ssp. female
genitalia. 30.
C. rothei
sp. n.
,
paratype, prep. PGy3738; 31.
C. dardistana
, prep. PGy3702; 32.
C. helix helix
, prep. PGy3697; 33.
C. irrisoria irrisoria
, prep. PGy3756.
Male genitalia. The genitalia of
C. rothei
(
Figs 14-17
) correspond closely to those of its congeners, especially
C. dardistana
(
Figs 18, 19
,
31
). The best keys for distinction are in the asymmetric saccular extensions. However, the large, long right saccular extension in
C. rothei
is distinctively longer than in other species with the exceptions of
C. dardistana
and
C. helix
(
Figs 20, 21
), while the everted vesica indicates slight differences in the size of the diverticuli and position of the cornutus.
C. rothei
can be separated from
C. dardistana
by the following: its somewhat more spatulate uncus; ventrally somewhat broader, subtriangular juxta; stronger, longer, terminally less tapering right saccular extension; longer, stronger left saccular extension; larger distal-secunder vesica diverticulum (oppositely projected from the aedeagus); a medially rather basally situated cornutus of the main vesica tube. Compared to
C. helix helix
and
C. helix dubatolovi
(
Figs. 22 23
) the neck of the cucullus is longer and the juxta is broader ventrally and shorter dorsally. The asymmetric saccular extensions are not acutely twisted as in
C. helix
, however, the right extension is nearly straight and more robust, whereas the left extension is straight, somewhat shorter, less elongate distally but acute terminally. The proximal-secunder diverticulum of the vesica (oppositely projected from the ductus ejaculatorius being slightly recurved to the dorsal part of aedeagus) is shorter than in
C. helix
.
Female genitalia.
C. rothei
(
Fig. 30
) has longer apophyses posteriores, a less calycular antrum and a smaller u-shaped medial depression in the ostium bursae than in
C. dardistana
(
Fig. 31
). In
C. helix
(
Fig. 32
) the antrum is less robust, the small medial depression in the ostium bursae is more vshaped and the appendix bursae longer.
C. irrisoria irrisoria
(
Fig. 33
) and
C. irrisoria nigrescens
have larger and subglobular corpus bursae.
Biology and distribution.
Long series of both sexes were collected at ultraviolet light between
7 and 12 June
2013
in west China's
Xinjiang Province
. It is likely endemic there in a remote area at the southwestern edge of the Taklimakan desert. Most specimens were collected in the Yarkan He river (
Fig. 13
) floodplain in leafy forest (tugay) dominated by Turanga Poplar, Oleaster (
Populus
,
Elaeagnus
) and
Salix
species
but some were also collected near sand dunes sparsely covered with Oleaster (
Elaeagnus
), Camel Thorn (
Alhagi
) and various
Zygophyllaceae (Caltrop)
bushes. Other desert
Noctuidae
species collected with
C. rothei
included
Catocala contemnenda
Staudinger
,
Catocala remissa
Staudinger
,
Drasteria antiqua
(Staudinger)
among others.
Etymology
.
The new species is named after a prominent German lepidopterist, Mr. Stefen Rothe (Taucha,
Germany
).