The family Onychiuridae (Collembola) from karst caves of the Basque biospeleologic district, with description of four new species Author Beruete, Enrique 0000-0002-3240-8946 University of Navarra, Faculty of Sciences, Department of Environmental Biology, University Campus, 31080 Pamplona, Spain. eberuete @ yahoo. es; https: // orcid. org / 0000 - 0002 - 3240 - 8946 eberuete@yahoo.es Author Arbea, Javier I. c / Ría de Solía 3, ch. 39. 39610 El Astillero (Cantabria). Author Baquero, Enrique 0000-0002-3240-8946 University of Navarra, Faculty of Sciences, Department of Environmental Biology, University Campus, 31080 Pamplona, Spain. eberuete @ yahoo. es; https: // orcid. org / 0000 - 0002 - 3240 - 8946 & University of Navarra, Institute of Biodiversity and Environment (BIOMA), University Campus, 31080, Pamplona, Spain * corresponding author: ebaquero @ unav. es; https: // orcid. org / 0000 - 0002 - 2145 - 8606 eberuete@yahoo.es Author Jordana, Rafael 0000-0002-3240-8946 University of Navarra, Faculty of Sciences, Department of Environmental Biology, University Campus, 31080 Pamplona, Spain. eberuete @ yahoo. es; https: // orcid. org / 0000 - 0002 - 3240 - 8946 eberuete@yahoo.es text Zootaxa 2021 2021-09-21 5040 2 151 194 journal article 10.11646/zootaxa.5040.2.1 1175-5326 5531192 CC1289B1-4FF9-4369-A596-9FF260C8F314 Protaphorura armata ( Tullberg, 1869 ) Lipura armata Tullberg, 1869 . Skand. Podurider Unterf. Lipurinae : 18 Onychiurus armatus (Tullberg) : Börner, 1902 . Zool. Anz ., 25(662): 97 Protaphorura armata (Tullberg) : Salt, Hollick, Raw & Brian, 1948 . J. Anim. Ecol ., 17(2): 145 Studied Material. Loc. 5. 1 ex. , 2.X.1987 . Loc. 17. 3 ex. , 15.VIII.1929 (MNCN_Ent 89461). Euedaphic species, probably cosmopolitan ( Jordana et al. 1997 ). It is found throughout the Iberian Peninsula. It is located both in forests and in meadows. The Arkalde cave specimen in Navarra was captured in beech litter in the debris cone under the mouth of the cavity ( Beruete 2000a , Beruete et al. 2002 ). It is a very common species in Navarra ( Jordana et al. 1997 ) so it can appear when dragged from outside. It has also been cited from the San Adrián cave in Gipuzkoa ( Stach 1954 ).