<document id="02C312E9B38758117B9D2FAB5E425ADB" ID-DOI="10.5252/zoosystema2024v46a2" ID-ISSN="1638-9387" ID-ZooBank="4B49C1D9-1196-4942-969F-2E923B1FC12C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1707561373589" checkinUser="plazi" docAuthor="Cupello, Mario" docDate="2024" docId="03C5B216FFEDFFB2DE85F9A86F9DF86B" docLanguage="en" docName="zoosystema.2024.46.2.pdf" docOrigin="Zoosystema 46 (2)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a2.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Gromphas , Dejean 1836" docType="treatment" docVersion="2" lastPageNumber="32" masterDocId="FFFCCA6EFFE6FFB9DD69FFFA6C63FF8D" masterDocTitle="The genus Gromphas Dejean, 1836 (Coleoptera, Scarabaeinae): nomenclature, distribution, and conservation, including a contribution to the debate on electronic publications in zoology" masterLastPageNumber="59" masterPageNumber="23" pageNumber="32" updateTime="1707561895447" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="93D1C58E5BA366CC097D7F491C06334A">The genus Gromphas Dejean, 1836 (Coleoptera, Scarabaeinae): nomenclature, distribution, and conservation, including a contribution to the debate on electronic publications in zoology</mods:title>
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<mods:namePart id="B3BB28F183E03E50C688ACF0F439411F">Cupello, Mario</mods:namePart>
<mods:affiliation id="E3C7B406E057DFBEB7270726E33AFC33">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná (Brazil); and Department of Entomology, Texas A &amp; M University, College Station, TX 77843 (United States) mcupello @ hotmail. com</mods:affiliation>
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<mods:date id="9E706C346CA3BB6D928A23B0A3FBDD2E">2024</mods:date>
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<emphasis id="B918DF12FFEDFFB2DE85F9A86835F9E1" box="[1004,1110,1618,1644]" italics="true" pageId="11" pageNumber="32">Gromphas</emphasis>
<bibRefCitation id="EFFD7EF1FFEDFFB2D934F9A86893F9E1" author="DEJEAN P. F. M. A." box="[1117,1264,1618,1644]" pageId="11" pageNumber="32" pagination="1 - 384" refId="ref31341" refString="DEJEAN P. F. M. A. 1836. - [livraisons 1 - 4: 1 - 384], in DEJEAN P. F. M. A. 1836 - 1837. Catalogue des Coleopteres de la collection de M. le Comte Dejean. Troisieme edition, revue, corrigee et augmentee. Mequignon-Marvis Pere et Fils, Paris, v-xiv + 503 p. [publication date for nomenclatural purposes: 30 th July 1836 (see Bousquet 2016)]" type="book chapter" year="1836">Dejean, 1836</bibRefCitation>
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<emphasis id="B918DF12FFEDFFB2DE44F96B6FEEF924" box="[813,909,1681,1705]" italics="true" pageId="11" pageNumber="32">Gromphas</emphasis>
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[
<date id="FFD225C0FFEDFFB2D93EF96B6892F924" box="[1111,1265,1681,1705]" pageId="11" pageNumber="32" value="1836-07-30">30th July 1836</date>
].
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. — 
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<emphasis id="B918DF12FFEDFFB2DE88F93F68CCF950" box="[993,1199,1733,1757]" italics="true" pageId="11" pageNumber="32">Coprobius lacordairii</emphasis>
Oken, 1834
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, by original monotypy.
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. — From the Latinisation of the Ancient Greek noun ΓΡΟμΦΆς (
<emphasis id="B918DF12FFEDFFB2DEEFF8D56FAEF8CA" box="[902,973,1839,1863]" italics="true" pageId="11" pageNumber="32">gromfás</emphasis>
), meaning ‘an old sow’, feminine in gender (
<bibRefCitation id="EFFD7EF1FFEDFFB2D81DF8D56F03F8EF" author="BRULLE A." pageId="11" pageNumber="32" pagination="19" refId="ref29964" refString="BRULLE A. 1838. - [livraison 8: p. 225 - 448], in BRULLE A. 1837 - 1838. Histoire naturelle des insectes, traitant de leur organisation et de leurs moeurs en general, par M. V. Audouin, et comprenant leur classification et la description des especes, par M. A. Brulle. Le tout accompagne de planches gravees sur acier, d'apres des peintures executees pour cette edition sur la collection du Museum de Paris. Tome VI. Coleopteres III. F. D. Pillot, Paris, 448 p. + pls. 18, 19, 19 bis, 20, 25 [publication date for nomenclatural purposes: 31 st of December 1838 (based on Bousquet 2016)] https: // www. biodiversitylibrary. org / page / 25546370" type="book chapter" year="1838">Brullé 1838</bibRefCitation>
; 
<bibRefCitation id="EFFD7EF1FFEDFFB2DE07F8B06F9AF8EF" author="HAROLD E." box="[878,1017,1866,1890]" pageId="11" pageNumber="32" refId="ref32645" refString="HAROLD E. 1869 a. - Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich: 979 - 1346 + [8]. [publication date for nomenclatural purposes: 27 th May 1969 (based on Bousquet 2016)] https: // www. biodiversitylibrary. org / page / 9641549" type="journal volume" year="1869">Harold 1869a</bibRefCitation>
; 
<bibRefCitation id="EFFD7EF1FFEDFFB2D961F8B06819F8EF" author="BAILLY A." box="[1032,1146,1866,1890]" pageId="11" pageNumber="32" refId="ref28925" refString="BAILLY A. 1895. - Dictionnaire grec-francais. Hachette, Paris, 2193 p." type="book" year="1895">Bailly 1895</bibRefCitation>
; 
<bibRefCitation id="EFFD7EF1FFEDFFB2D9E1F8B06903F8EF" author="LIDDELL H. G. &amp; SCOTT R." box="[1160,1376,1866,1890]" pageId="11" pageNumber="32" refId="ref33789" refString="LIDDELL H. G. &amp; SCOTT R. 1897. - A Greek-English lexicon. Revised and augmented throughout with the cooperation of professor Drisler, of Columbia College, New York. Eighth Edition. Harper &amp; Brothers, New York, New York, xvi + 1778 p." type="book" year="1897">Liddell &amp; Scott 1897</bibRefCitation>
; 
<bibRefCitation id="EFFD7EF1FFEDFFB2D807F8B06F03F8F1" author="BROWN R. W." pageId="11" pageNumber="32" refId="ref29817" refString="BROWN R. W. 1954. - Composition of scientific words. A manual of methods and a lexicon of materials for the practice of logotechnics. Smithsonian Books, Washington, D. C., 882 p. https: // archive. org / details / compositionofsci 00 brow" type="book" year="1954">Brown 1954</bibRefCitation>
; 
<bibRefCitation id="EFFD7EF1FFEDFFB2DE06F89E68C7F8F1" author="CUPELLO M. &amp; VAZ-DE-MELLO F. Z." box="[879,1188,1892,1916]" pageId="11" pageNumber="32" pagination="439 - 482" refId="ref30610" refString="CUPELLO M. &amp; VAZ-DE-MELLO F. Z. 2013. - Taxonomic revision of the South American dung beetle genus Gromphas Brulle, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina). Zootaxa 3722 (4): 439 - 482. https: // doi. org / 10.11646 / zootaxa. 3722.4.2" type="journal article" year="2013">Cupello &amp; Vaz-de-Mello 2013</bibRefCitation>
). The name likely draws a parallel between the stout, hornless body shape of both pigs and the only species originally included in the genus, 
<taxonomicName id="4C6C7883FFEDFFB2D982F8606908F83C" authorityName="Oken" authorityYear="1834" box="[1259,1387,1945,1969]" class="Insecta" family="Scarabaeidae" genus="Coprobius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="32" phylum="Arthropoda" rank="species" species="lacordairii">
<emphasis id="B918DF12FFEDFFB2D982F8606908F83C" box="[1259,1387,1945,1969]" italics="true" pageId="11" pageNumber="32">G. lacordairii</emphasis>
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, while also making a probable reference to the ‘filthy’ coprophagous habits of these dung beetles.
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