Identity of species-group taxa of the Western Palaearctic Clytrini (Coleoptera: Chrysomelidae) described by Maurice Pic and Louis Kocher
Author
Bezdċk, Jan
Author
Regalin, Renato
text
Acta Entomologica Musei Nationalis Pragae
2015
suppl.
2015-10-15
55
1
114
journal article
9543
10.5281/zenodo.4272771
77c874cd-4f85-4746-8466-3ca09e2c2b8d
0374-1036
4272771
urn:lsid:zoobank.org:pub:A431893C-11D8-4480-ABE1-BB65A823C734
Clytra
(
Clytraria
)
atraphaxidis atraphaxidis
(
Pallas, 1773
)
Chrysomela atraphaxidis
Pallas, 1773: 725
(original description).
Clythra atraphascidis
[sic!] var.
delagrangei
Pic, 1896a: 30
(nomen nudum).
Elythra
[sic!]
atraphaxidis
var.
delagrangei
Pic, 1896b: 62
(original description).
Clytra atraphaxidis
var.
nigromaculata
Pic, 1897a: 84
(original description),
syn. nov.
Clytra atraphaxidis
var.
quinquemaculata
Pic, 1920a: 7
(unavailable infrasubspeciħc name).
Clytra atraphaxidis
var.
milliati
Pic, 1942b: 7
(original description).
Type
localities.
Chrysomela atraphaxidis
: ‘Irtin’ [= western Siberia/
Kazakhstan
, Irtysh river].
Clythra atraphaxidis
var.
delagrangei
: ‘
Akbès’
[=
Turkey
near the city of
ø
skenderun
].
Clytra atraphaxidis
var.
nigromaculata
: ‘Bakou’ [=
Azerbaijan
,
Baku
].
Clytra atraphaxidis
var.
quinquemaculata
: ‘Turkestan et environs de Smyrne’.
Clytra atraphaxidis
var.
milliati
: ‘Liban’ [=
Lebanon
].
Type
material examined.
Chrysomela atraphaxidis
: not examined.
Clythra atraphaxidis
var.
delagrangei
:
SYNTYPE
: 1 ♂, ‘301 [w, h] // type. [w, h] // Delagrangei [w, h] // Akbès / n sp.? [w, h] //
Delagrangei Pic
[w, h] // Titubaea / arabica / Akbes 94 [w, h] //
TYPE
[r, p] // v.
Delagrangei Pic
[grey label, h] // Museum Paris / Coll. M. Pic [w, p]’ (
MNHN
– coll. generale).
Clytra atraphaxidis
var.
nigromaculata
:
SYNTYPES
: 1♂, ‘type. [w, h] // atraphascidis /
Baku
[w, h] //
v.nigromaculatus
/ Pic [w, h] //
TYPE
[r, p] //
v. nigromaculata Pic
[grey label, h] // Museum Paris / Coll. M. Pic [w, p]’ (
MNHN
– coll. generale);
1 ♂, ‘type. [w, h] //
Baku
[w, h] //
Clythra
/ atraphaxidis [w, h] // atrapascidis /
v. nigromaculatus
[w, h] //
TYPE
[r, p] // Museum Paris / Coll. M. Pic [w, p]’ (
MNHN
– coll. generale).
Clytra atraphaxidis
var.
milliati
:
SYNTYPE
:
1 ♀
, ‘M… /
Liban
[partly illegible, w, h] // ex Milliat [w, h] // v. Milliati / mihi [w, h]’ (
MNHN
– coll. Pic).
Original material of infrasubspeciħc entity.
Clytra atraphaxidis
var.
quinquemaculata
: 1 ♂, ‘
type
[w, h] // v. 5 maculata / Pic [w, h] // Turkestan / Aulie-Ata / C.
ARIS
[w, p] // v. 5 maculata Pic [grey label, h] //
TYPE
[r, p] // Museum Paris / Coll. M. Pic [w, p]’ (
MNHN
– coll. generale).
Distribution.
South Europe, Caucasus, Near East, Central Asia (for details see
REGALIN & MEDVEDEV 2010b
).
Comments.
PIC (1896b
,
1897a
,
1920a
,
1942b
) described four varieties of
Clytra atraphaxidis
. Two of them,
var.
delagrangei
and
var.
milliati
, are correctly treated as colour aberrations and synonyms of
C. atraphaxidis atraphaxidis
(e.g.
REGALIN & MEDVEDEV 2010b
). The additional two,
var.
nigromaculata
and
var.
quinquemaculata
, were listed as aberrations of
C. atraphaxidis
ssp.
maculifrons
Zubkov, 1833
by
WINKLER (1929)
, which was followed also by
REGALIN & MEDVEDEV (2010b)
.
Clytra atraphaxidis
ssp.
maculifrons
differs from the nominotypical subspecies in the head orange or orange with black spot on vertex and is distributed in Central Asia southernmost to
Iran
and
Afghanistan
and westernmost to Caucasus (REGALIN & MED- VEDEV 2010b). Because
var.
nigromaculata
has head black as the nominotypical subspecies we transfer it as a new synonym of
C. atraphaxidis atraphaxidis
.
It is necessary to note that we have some doubts about the validity of
ssp.
maculifrons
.
Clytra atraphaxidis
is a widely distributed species with very variable dorsal pattern. Nominotypical subspecies and
ssp.
maculifrons
can be distinguished only by the colouration of the head, which, in view of colour variability throughout the whole range, seems to be an insufħcient character to separate the subspecies. However, we leave this problem for future studies.
Clytra atraphaxidis
var.
quinquemaculata
was described in the paper containing also description of subspecies (
PIC 1920a
), thus became infrasubspeciħc according to Article 45.6.4 of the Code (
ICZN 1999
). The original specimen of this variety is conspeciħc with
Clytra atraphaxidis atraphaxidis
as it has a black head.
Clythra atraphaxidis
var.
delagrangei
was ħrst mentioned by
PIC (1896a)
without a relevant description, and we treat it as a nomen nudum. The correct description was published by
PIC (1896b)
.